Lesbosoxylon kemaliyensis Akkemik & Mantzouka, 2020

Akkemik, Ünal, Mantzouka, Dimitra & Yildirim, Demet Kiran, 2020, The first report of Lesbosoxylon Süss & Velitzelos from the early-middle Miocene of eastern Anatolia, Geodiversitas 42 (23), pp. 427-441 : 432

publication ID

https://doi.org/ 10.5252/geodiversitas2020v42a23

publication LSID

urn:lsid:zoobank.org:pub:54A81360-9A19-456B-AF72-E4868A4835BA

DOI

https://doi.org/10.5281/zenodo.4488612

persistent identifier

https://treatment.plazi.org/id/03A987CE-FFF8-0208-83AC-FC05FABDFC91

treatment provided by

Felipe

scientific name

Lesbosoxylon kemaliyensis Akkemik & Mantzouka
status

sp. nov.

Lesbosoxylon kemaliyensis Akkemik & Mantzouka , sp. nov. ( Figs 3-6 View FIG View FIG View FIG View FIG )

Plant Fossil Names Registry Number: PFN001780

HOLOTYPE. — Specimen KEM01 About KEM ; repository: Istanbul University-Cerrahpasa, Faculty of Forestry, Department of Forest Botany (designated here).

ETYMOLOGY. — The epithet “ kemaliyensis ” comes from the name of the main city “Kemaliye” nearby the fossil site.

TYPE HORIZON. — Divriği Formation.

AGE. — Early-middle Miocene.

TYPE LOCALITY. — Kemaliye (Eğin), Erzincan, Turkey.

DIAGNOSIS. — Coniferous wood with resin canals. Growth ring boundaries distinct, transition from earlywood to latewood mostly gradual, axial and radial resin canals with thin-walled epithelial cells, latewood tracheids thin to thick walled, bordered pits on radial walls of tracheids 1-2(-3) seriate, crassulae common, rays heterocellular, uniseriate, partly biseriate, uniseriate rays up to 27 cells high, fusiform rays up to 30 seriate high. Axial parenchyma occasionally present. Ray tracheids 2-3 rows, cell walls of ray tracheids smooth. Cross-field pitting pinoid, 1-2(-6) pits per cross-field, horizontal and end walls of ray parenchyma cells smooth and slightly nodular, horizontal walls occasionally pitted, indentures not observed.

DESCRIPTION

The petrified wood sample has grey color; dimensions are 7 × 8 × 19 cm ( Fig.3 View FIG ). In cross section,growth ring boundaries are distinct, transition from earlywood to latewood is mostly gradual ( Fig.4A, B View FIG ). Measured diameters of tracheids and pits are given inTable 1. Axial resin canals with thin walled epithelial cells present mostly in latewood zone ( Fig. 4A, B View FIG ). Traumatic resin canals present ( Fig. 4C View FIG ). In tangential section, rays uniseriate ( Fig.4D View FIG ), and fusiform rays present ( Fig. 4E View FIG ). Rays occasionally biseri-ate or partly biseriate ( Fig. 5A View FIG ). Big rays with idioblast-like cells present ( Fig 5 View FIG B-D). Ray cells uniform in some rays, and heterocellular, wider or narrower ( Fig. 5 View FIG D-E). Height of uniseriate rays up to 27 cells ( Fig. 4C View FIG ), height of fusiform rays up to 30 cells and wider ( Fig. 4D View FIG and Fig. 5D View FIG ). Average height of rays 4-15 cells. Radial resin canals with thin–walled epithelial cells. Axial parenchyma cells occasionally present, and generally smooth transverse end walls ( Fig. 5 View FIG F-G and I), and slightly nodular ( Fig. 5H View FIG ). Axial resin canal visible in radial section ( Fig. 6A View FIG ). Tracheid pitting in radial walls of earlywood 1-2 (-3) seriate opposite and crassulae formation common ( Fig. 5 View FIG B-D). Cross-field pitting pinoid in 1(-2) rows of usually 1-2(-6) pits per cross-field ( Fig. 6E, F View FIG ). End walls of ray parenchyma cells smooth, no indentures observed. Rays heterocellular. Ray tracheids present, 1-3 seriate, smooth walled ( Fig. 6G View FIG ). Ray cells smooth walled ( Fig. 6 View FIG E-H) and occasionally pitted. These descriptive features are also provided in Appendix A based on “Criteria used according the IAWA for fossil conifers” by Wheeler et al. (2004).

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