Uroxys latesulcatus Bates, 1891

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 242-243

Uroxys latesulcatus Bates, 1891 Plates 53C, 58 A–B

Uroxys latesulcatus Bates, 1891: 24 (original description. Type locality: Ecuador: Pichincha, 12,000 feet [= 3655 m]; Machachi, 9-10,000 feet [= 2745-3050 m]).

Uroxys latesulcatus : Gillet 1911a: 50 (catalog); Campos 1921: 55 (cited for Ecuador); Arrow 1933: 388 (characters in key); Vulcano and Pereira 1967: 580 (characters in key); Carvajal et al. 2011: 318-319 (cited for Ecuador); Krajcik 2012: 262 (complete list of species); Chamorro et al. 2018: 98 (cited for Ecuador).

Uroxys latesulcata : Blackwelder 1944: 203 (list of species for Latin America).

Uroxys magnus Balthasar, 1940: 37 (original description); Vulcano and Pereira 1967: 580 (characters in key); Carvajal et al. 2011: 318-319 (cited for Ecuador); Bezdek and Hajek 2011: 359 (catalog of types NMPC); Krajcik 2012: 262 (complete list of species, cited as Uroxys magnus Balthasar, 1947), Chamorro et al. 2018: 98 (cited for Ecuador), syn. n.

Type specimens.

Uroxys latesulcatus Bates, 1891. The lectotype (♂) (here designated) and two paralectotypes are deposited at the MNHN (ex coll. HW Bates and ex coll. R Oberthur). Locality: Ecuador (examined). One paralectotype is deposited at the NHML, examined.

Lectotype (here designated) (♂): "Ecuador. / feet. / Ed. Whymper. [p]", "Uroxys / latesulcatus / Bates [p, black margin]", “Ex-Musæo / H. W. BATES / 1892 [p, black margin]", “MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [p, green label, black margin]", "LECTOTYPE ♂ / Uroxys / latesulcatus / Bates / des. F.Z. Vaz-de-Mello. 2014 [p and hw, red label, black margin]".

Paralectotype (♂): "Ecuador. / feet. / Ed. Whymper. [p]", "Uroxys / latesulcatus / Bates [p, black margin]", “Ex-Musæo / H. W. BATES / 1892 [p, black margin]", “MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [p, green label, black margin]", "PARALECTOTYPE / Uroxys ♂ / latesulcatus / Bates / des. F.Z. Vaz-de-Mello, 2014 [hw and p, yellow label, black margin]".

Paralectotype (♂): "Ecuador. / feet. / Ed. Whymper. [p]", "latesulcatus / Bates [p]", “Ex-Musæo / H. W. BATES / 1892 [p, black margin]", “MUSÉUM PARIS / 1952 / COLL. R. OBERTHUR [p, green label, black margin]", "PARALECTOTYPE / Uroxys ♂ / latesulcatus / Bates / des. F.Z. Vaz-de-Mello, 2014 [hw and p, yellow label, black margin]".

Paralectotype (♂): "Uroxys / latesulcatus, / (Type) Bates [hw]", "92-24 [p]", "SYN- / TYPE [p, blue label]", "PARALECTOTYPE / Uroxys ♂ / latesulcatus / Bates / des. F.Z. Vaz-de-Mello, 2014 [hw and p, yellow label, black margin]".

Uroxys magnus Balthasar, 1940. The lectotype (♂) (here designated) is deposited at the NMPC (ex coll. V Balthasar). Locality: Ecuador (examined).

Lectotype (here designated) (♂): "Ecuador / D. Stübel / 5351 [hw, green label]", "4684 [hw, red letters]", "Typus [p, red label]", "Uroxys / magnus / n. sp. / DR. BALTHASAR. DET. [p]", "magnus / m. [hw, green label]", "Mus. Nat. Pragae / 65711 / Inv. [p and hw, red label]", "LECTOTYPE ♂ / Uroxys / magnus / Balth / des. F.Z. Vaz-de-Mello. 2013 [p and hw, red label, black margin]".

Distribution.

Only known from Ecuador.

Records examined.

BOLIVAR: Cashca Totoras (5 specimens MQCAZ). COTOPAXI: Sigchos (2 specimens MQCAZ). IMBABURA: Selva Alegre (3 specimens MQCAZ). PICHINCHA: Atahualpa, Bosque Protector Piganta, 2880 m (2 specimens MUTPL); Cochasquí, 3100 m (56 specimens CEMT); Chiriboga (5 specimens MQCAZ); EL Chalpar, 3300 m (1 specimen CEMT, 3 specimens MQCAZ); Machachi, 2700 m (21 specimens MQCAZ); La Cocha (1 specimen CEMT; 8 specimens MQCAZ); Nono (8 specimens MQCAZ); Palmeras, 2200 m (5 specimens MQCAZ); Pasochoa, 3500 m (1 specimen CEMT; 8 specimens MQCAZ); Pingtag, 2880 m; Quito, Cumbayá, 2340 m (13 specimens MQCAZ); Reserva Ecológica Los Illinizas (2 specimens MUTPL); San José de Minas (3 specimens CEMT; 4 specimens MQCAZ); Yanacocha (2 specimens MQCAZ). UNDETERMINED PROVINCE: without specific locality (3 specimens MNHN; 1 specimen NHML, 1 specimen NNPC).

Literature records.

PICHINCHA: without specific locality 12.000 feet [= 3655 m] ( Bates 1891: 24); Machachi 9-10000 feet [= 3045 m] ( Bates 1891: 24).

Temporal data.

Collected in January, February, March, April, May, June, October, November, and December.

Remarks.

Inhabits the high montane evergreen forests of the Andean region from 2340-3300 m a.s.l. Collected with pitfall trap baited with pig or cow feces.

Balthasar in 1940, described Uroxys magnus for Ecuador (without type locality) as a different species from Uroxys latesulcatus Bates, 1891 (recorded for Ecuador, Pichincha at 12,000 feet [= Pichincha volcano] and Machachi 9 at 10,000 feet, as type localities). However, upon examining the external and genital morphology of the type specimens, U. latesulcatus (lectotype ♂ here designated, deposited at the MNHN, Plate 58A) and U. magnus (lectotype ♂ here designated, deposited at the NMPC, Plate 58B), we believe that they belong to the same species due to similar characteristics (specifically, the shape of the head, dorsal colouration, body length and aedeagus). Therefore, we propose that U. magnus is a synonym of U. latesulcatus (see ICZN 1999, Article 23). Two lectotypes (without specific localities for Ecuador) are here designated and illustrated (♂ Plate 58A, B).