Kleinzaches germanicianus, Alekseev & Tomaszewska, 2018

Alekseev, Vitalii I. & Tomaszewska, Wioletta, 2018, New handsome fungus beetles (Coleoptera: Coccinelloidea: Anamorphidae, Endomychidae) from European amber of the Upper Eocene, Palaeontologia Electronica (6 A) 21 (1), pp. 1-23 : 5-7

Kleinzaches germanicianus sp. nov.

Figure 2

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Type material. Holotype: No. 181-1 [ CCHH]. Sex unknown. The beetle inclusion is preserved in a polished piece of transparent amber, yellowish in color. The small amber piece is embedded in a block of polyester resin (total measurements are 9 x 8 x 3 mm). Syninclusions consist of numerous small wood pieces and three gas bubbles. The right hind leg and terminal antennomere of right antenna of the beetle are missing.

Type strata. Bitterfeld amber. Eocene.

Type locality. Germany, Sachsen-Anhalt, Goitzsche (Bitterfeld-Wolfen) .

Etymology. The specific epithet refers to the region of the specimen’s origin and deposition: “germanicianus ” in Latin literally means “stationed in Germany.”

Diagnosis. As stated for new genus.

Description. Body length = 1.45 mm; width (at widest point in middle of elytra) = 1.25 mm. Elytral length (along suture) = 1.05 mm. Habitus: broadly oval, dorsally convex; dorsal surface shiny, covered with long erect hairs; body uniformly dark brown with appendages and ventrites 2–5 lighter.

Head retracted into prothorax up to position of compound eyes, pubescent. Compound eyes rounded, weakly prominent. Fronto-clypeal suture distinct. Antennae composed of 10 antennomeres, extending to half length of pronotum, with loose club composed of three antennomeres. Antennomere 8 oblong, almost twice as long as wide; antennomere 9 as long as wide; antennomere 10 symmetrical, oblong and pointed. Terminal labial and maxillary palpomeres oblong, weakly pointed.

Pronotum transverse (2.5 times as wide as long), widest within one-third of its length; bordered laterally and anteriorly. Lateral sulci long, narrow, spanning posteriormost three-fourths of pronotal length. Pronotum covered with sparse, irregular, semirecumbent pubescence. Pronotal posterior margin almost straight. Procoxal cavities externally open, internally closed. Prosternal process somewhat triangular with rounded angles, distinctly bordered throughout. Mesocoxal cavities widely closed laterally.

Elytra rounded, convex, with base slightly wider than posterior margin of pronotum, covered with long pubescence, with small strial punctures, arranged in almost regular rows. Distance between punctures in each row 1.0–2.0 times larger than diameter of each point, and intersticial width is about three times diameter. Spaces between rows with regular long erect setae, separated from each other by distance shorter than length of each seta. Elytral epipleura long, reaching ventrite 5; with rows of small punctures at outer margin.

Abdomen with five ventrites; apex of terminal ventrite widely rounded; postcoxal lines absent. Ventrite length proportions (measured along midline) according to formula: 17: 6: 6: 6: 11. Metacoxae widely separated by intercoxal process of ventrite 1; process is twice as wide as longitudinal metacoxal diameter and straight anteriorly. Distance between irregular and fine punctures on abdominal ventrites is 1.0–3.0 times larger than diameter of each puncture.

Legs with femora thickened apically. Tibiae slender. Tarsi long, with simple cylindrical segments. Tarsal formula 3-3-4. Metatarsomeres 1–3 almost equal in length; metatarsomere 4 longest, almost as long as tarsomeres 1–3 combined. Tarsal claws quite long (0.3 times as long as apical tarsomere), symmetrical, appendiculate.

Remarks. The periscutellar area (scutellum, central part of pronotal base and basal part of elytra) is invisible because of gas bubbles.

Figures

FIGURE 2. Kleinzaches germanicianus gen. et sp. nov. (No. 181-1 [CCHH], holotype) (Anamorphidae) from Bitterfeld amber. 1-2, Habitus in dorsal (1) and ventral (2) views. 3, A reconstruction of the habitus in dorsal view. 4, Close-up of the anterior part of body in latero-ventral view.