Iteaphila sicamous Sinclair, 2012

Sinclair, Bradley J. & Shamshev, Igor V., 2012, 3561, Zootaxa 3561, pp. 1-61 : 45

Iteaphila sicamous Sinclair sp. nov.

( Figs. 11C, 14B)

Type material. HOLOTYPE, ♂ labelled: “ 8 km E. Sicamous, B.C. [ CANADA: Bristish Columbia ]/ 8-VI-1992 / A. Borkent CD1419”; “ HOLOTYPE / Iteaphila / sicamous/ Sinclair [red label]” ( CNC) . PARATYPE: same data as holotype (1 ♂, CNC) .

Recognition. This species is distinguished by its very long proboscis, 1.5 times head height; biserial acrostichal setae; abdominal setae pale; apex of postgonites slender, sinuous and lacking setae or pubescence; cercus short and slender and hypoproct process absent.

Description. Wing length 3.7–4.4 mm. Male. Head subcircular, black, with black to pale setation; occiput brownish pollinose. Holoptic, with upper ommatidia enlarged. Frons represented by small triangular space just above antennae, bare, brownish grey pollinose. Ocellar triangle very prominent, with pair of long, thin, proclinate ocellar setae. Postvertical and postocular setae long, thin, hair-like; occiput covered with numerous similar setae longer on lower part and generally paler. Antenna brown; scape short, subequal to globular pedicel, both with short setae; postpedicel nearly 6 times longer than basal width, smoothly tapered; stylus very short, shorter than basal width of postpedicel, bristle-tipped, length of segment 9 slightly shorter than width, apical bristle longer than segment 9. Proboscis very long, 1.5 times head height, projected obliquely; labium longer than head height; palpus projected parallel to labrum, two-thirds length of labrum.

Thorax black (postpronotal lobe yellowish posteriorly and postalar ridge brownish yellow), with black setation; smoothly arched, prescutellar depression very slightly prominent; scutum viewed dorsally largely velvety brown, somewhat shiny, lacking vittae. Proepisternum with several brown setae in lower and upper parts. Postpronotal lobe with several fine setae and 1 prominent seta. Mesonotal setae mostly prominent, 4 npl, 2–3 prescutellar dc, 1 pal and sctl somewhat stronger; acr biserial, long, separated by less than half length of setula, ending at prescutellar depression; dc long, subequal to acr, uniserial; 6 pairs of sctl in single row.

Legs yellowish brown. Coxae and trochanters with several short, fine setae. Fore and mid femora with row of long posteroventral setae. Hind femur with row of long anteroventral and dorsal and short posteroventral hair-like setae. Tibiae lacking prominent setae. Tarsomere 5 on all legs slightly flattened; pulvilli broad, shorter than tarsal claw.

Wing infuscate, with brown veins; no bristle at wing base; stigma distinct brownish, elliptical, overlapping apex of vein R 1; anal lobe very prominent, acute. R 2+3 slightly arched beneath stigma; radial fork with base well proximal to apex of R 2+3; R 5 and M 1 slightly divergent before wing-apex; cell d broad, longer than basal cells, truncate apically; m-m crossvein long, M branches widely separated; dm-cu crossvein slightly curved; M 4 shorter to longer than dm-cu crossvein. Apex of cell cua slightly rounded, CuP+CuA long, ending just short of wing margin. Halter dark, base of stem light brown.

Abdomen narrowed proximad, covered with numerous, pale, long hair-like setae, posteromarginal setae not prominent; tergites viewed dorsally subshiny, finely pollinose, viewed posteriorly brownish pollinose; sternites finely pollinose. Terminalia concolorous with abdomen, enlarged. Hypandrium robustly rounded, upcurved apically with broad truncate apex; clothed in short sparse setae, with some longer posterior setae; gonocoxal apodeme long and broad. Postgonite posterior to phallus, extended posteriorly beyond epandrium, parallel with phallus and strongly bent nearly at right angles subapically; bent apical section sinuous, long and slender fused with bifid tip, lacking microtrichia. Epandrium not greatly inflated laterally; dorsal bridge narrow; anterior portion subquadrate and very broad, posterior third greatly narrowed compared to anterior portion; inner face of apex of surstylus unmodified. Phallus long and robust, arched beyond epandrium posteriorly, strongly bent subapically; apical section broad and straight; apex truncate and partially membranous; ejaculatory apodeme plate-like, rounded, longer than gonocoxal apodeme. Base of phallus produced posteriorly into trifid lobe; lateral pair sickleshaped, slender; median lobe triangular, with broad base. Cercus tapered, very short, less than one-third length of epandrium; finger-like with apex tapered, slightly curved, narrowly rounded; hypoproct not produced into paired lobes ( Fig. 11C).

Female. Unknown.

Etymology. The specific name is a noun in apposition, named after the type locality.

Distribution. This species is known from a single locality in southern British Columbia ( Canada) ( Fig. 14B).

Remarks. On the basis of the male terminalia (shape of cercus) and very long mouthparts, this species is apparently closely related to I. triangula .