Iteaphila rasnitsyni Shamshev, 2012

Sinclair, Bradley J. & Shamshev, Igor V., 2012, 3561, Zootaxa 3561, pp. 1-61 : 42-43

Iteaphila rasnitsyni Shamshev sp. nov.

( Figs. 11A, 12A)

Type material. HOLOTYPE, ♂ labelled: “[ RUSSIA] Kedrovaya Pad' Nature Reserve / south Primor'e / Rasnitsyn 7.VI.1962 [In Russian]”; “ HOLOTYPE / Iteaphila / rasnitsyni / Shamshev [red label] [specimen in poor condition]” ( ZIN).

Recognition. This species is distinguished by the slender cercus, broad anvil-shaped phallic process and erect surstylus, projecting posterodorsally.

Description. Wing length 3.3 mm. Male. Head black in ground-colour, with brown to black setation, occiput finely greyish pollinose. Holoptic, with upper ommatidia enlarged. Frons represented by small triangular space just above antennae, bare, greyish brown pollinose. Ocellar triangle prominent, with 2 pairs of thin setae; anterior ocellars rather long, posterior ocellars short. Postvertical and postocular setae thin, moderately long; occiput covered with numerous short bristly hairs in lower part. Antenna brown; scape short, subequal to globular pedicel, both with short setae; postpedicel rather narrow, nearly 4.0 times longer than wide, smoothly tapered; stylus very short, bristle-tipped; segment 9 short, hardly distinguishable, apical bristle about as long as segment 9. Proboscis long, projected oblique; labium about half height of head; palpus projected parallel to labrum, nearly as long as labrum.

Thorax black in ground-colour, with black setation; scutum viewed dorsally dull brown, viewed anteriorly velvety brown; pleura uniformly greyish pollinose [damaged on prescutellar depression and scutellum]. Proepisternum with several brownish, short hairs in lower and upper parts. Postpronotal lobe with 1 long and several short setae. Mesonotal bristles prominent but thin; acr short, arranged in 2 close rows, lacking on prescutellar depression; dc 1–2-serial, offset from rows anteriorly, in anterior part somewhat longer than acr; several short presut and psut spal, 4 npl (with additional thinner and shorter setae).

Legs almost wholly brownish, knees yellowish, unmodified. Coxae and trochanters with ordinary bristly hairs. Fore femur with short, thin posteroventral and dorsal setae. Mid femur with similar pattern of setation but posteroventral setae longer and stronger on apical half. Hind femur with moderately long, thin, anteroventral and dorsal setae. Tibiae lacking prominent bristles. Tarsomere 5 on all legs flattened; pulvilli broad, shorter than tarsal claw.

Wing finely brownish infuscate, with brownish veins; no bristle at wing base; stigma distinct brownish, elliptical, overlapping apex of vein R 1; anal lobe very prominent, acute. R 2+3 slightly sinuate in apical part; radial fork V-shaped, with base proximal to apex of R 2+3; R 5 and M 1 somewhat divergent before wing-apex; cell d broad, longer than basal cells, not produced apically (truncate); m-m crossvein long, M branches widely separated; dm-cu crossvein straight; M 4 somewhat longer than dm-cu crossvein. Apex of cell cua slightly rounded, CuP+CuA long, ending short of wing margin. Halter brown.

Abdomen brown, subshining (from any view), finely greyish pollinose, covered with long pale bristly hairs. Terminalia concolorous with abdomen, small. Hypandrium slightly rounded, upcurved apically with broad truncate apex; gonocoxal apodeme expanded apically. Postgonite posterior to phallus, extended slightly beyond epandrium, parallel with phallus and strongly arched subapically; bent apical section subequal to apex of phallus, paired with apex membranous, bearing roughened surface and teeth-like projections, apex rounded. Epandrium quadrate, not greatly inflated laterally; dorsal bridge very narrow; apex of surstylus very slender, projecting posterodorsally. Phallus long and slender, arched beyond epandrium, strongly recurved subapically; apical section long; apex rounded and membranous; ejaculatory apodeme plate-like, rounded, longer than gonocoxal apodeme; base of phallus produced posteriorly into anvil-shaped lobe with pair of slender lateral prolongations ( Figs. 11A). Cercus tapered and slender, slightly longer than half length of epandrium, finger-like and broad basally, apex narrow rounded, extended free from epandrium; hypoproct produced into pair of long, divergent lobes, projecting free from cercus.

Female. Unknown.

Etymology. The species is dedicated to the Russian palaeontologist and entomologist Professor Alexander P. Rasnitsyn (Palaeontological Institute of Russian Academy of Sciences, Moscow).

Distribution. This species is known from a single locality in the southern region of the Russian Far East ( Fig. 12A).