Ipomoea funis Schldt. & Cham.

Wood, John R. I., Munoz-Rodriguez, Pablo, Williams, Bethany R. M. & Scotland, Robert W., 2020, A foundation monograph of Ipomoea (Convolvulaceae) in the New World, PhytoKeys 143, pp. 1-823 : 1

315. Ipomoea funis Schldt. & Cham. , Linnea 5: 118. 1830. (Schlechtendal and Chamisso 1830: 118)

Morenoa grandiflora La Llave in La Llave & Lex. , Nov. Veg. Descr. Fasc. 1: 17. 1824. (La Llave and Lexarza 1824: 17), non Ipomoea grandiflora (L.f.) Lam. (1791). Type. MEXICO. San José del Corral, sine col. (wherabouts unknown).

Quamoclit grandiflora (La Llave) G. Don , Gen. Hist. 4: 259. 1838. (Don 1838: 259).

Ipomoea llaveana Meisn. in Martius et al. , Fl. Brasil. 7: 219. 1869. (Meisner 1869: 219). Type. Based on Morenoa grandiflora La Llave

Quamoclit langlassei House , Bull. Torrey Bot. Club. 36; 597. 1909. (House 1909b; 597). Type. MEXICO. probably Guerrero, E. Langlassé 875 (holotype US00111497, isotypes GH, K, P).

Ipomoea funis var. langlassei (House) O'Donell , Lilloa 29: 41. 1959 ( O’Donell 1959a: 41).

Type.

MEXICO. Veracruz, San Andrés, Schiede & Deppe 556 (lectotype HAL98219, designated here; isolectotype NY).

Description.

Climbing herb, stems glabrous or pubescent. Leaves ovate, petiolate, 7-12(-17) × 3-8(-13) cm, entire ( var. langlassei ), undulate, irregularly dentate or 3-lobed, base cordate sometimes with a square sinus, apex finely acuminate, both surfaces thinly pubescent to glabrous; petioles 2.5-15 cm. Inflorescence of long-pedunculate few-flowered cymes; peduncles 10-35 cm; bracteoles 0.5-3 mm, ovate, caducous; secondary to ultimate peduncles 1-1.5 cm; pedicels 7-35 mm; sepals with scarious margins, outer 4-5 × 4 mm, ovate, obtuse or truncate, strongly mucronate, awn 3-9 mm, inner elliptic, slightly longer and broader with a similar awn; corolla 5-6 cm long, red, glabrous, funnel-shaped, basal cylindrical tube 2.5-3 cm, then strongly widened, limb broad, c. 5 cm diam., shallowly lobed, stamens weakly exserted. Capsules globose, glabrous; seeds tomentose with scattered tufts of longer hairs.

Illustration.

Figure 3H .

Distribution.

Endemic to central Mexico, where it grows in disturbed bushy places, especially in damp gullies and along streams between 1600 and 2300 m.

MEXICO. Dist. Fed.: A. García-M 4368 (MEXU) - possibly introduced. Guanajuato: fide O’Donell (1959a: 40). Guerrero: Mina, G.B. Hinton 9866 (K); ibid., Fresnos, 9752 (K); Leonardo Bravo, M. Castro 153 (IEB); J.C. Soto 7496 (MEXU). Michoacán: Aguililla, Y. Ramírez-Amezcua & V. Steinmann 1222 (ARIZ, IEB); Nuevo San Juan Parangaricutiro, V.W. Steinmann & J.M. Porter 3982 (IEB, MEXU); San Miguel, Leavenworth & Hoogstrahl 1075 (MO); Cerro Tancítaro, Apo, R. McVaugh 24874 (MICH). Oaxaca: H. Galeotti 1358 (K); Miahuatlán, T. Croat 46031 (MO); Santa Cruz Itundujia, K. Velasco-G. et al. 2823 (IEB); Sierra de Mihuatlán, F. Miranda 8836 (MEXU). Puebla: Jardín del Calvario, G. Arsène 2339 (BM, K, MO, NY, US). San Luis de Potosí: Verles d’Aoust 1882 (P) - location seems improbable. Veracruz: M. Botteri 465 (BM); Orizaba, E. Bourgeau 2985 (K, P); A. Barrera et al. 331 (MEXU); L.A. Castillo-Hernández et al. 342 (MEXU).

Typification.

We have designated the specimen at Halle as lectotype of Ipomoea funis as it has Schiede’s original label attached to the sheet.

Notes.

O’Donell recognised Ipomoea funis var. langlassei for plants recorded from Guerrero which differ from the type in their entire, adaxially nearly glabrous leaves. However, there is much variation overall in this species in leaf shape, lobing and dentation as well as in indumentum, and we do not think this variety merits recognition.

A specimen from Chinicuila ( Michoacán) J.C. Soto Nuñez et al. 11115 (MEXU) appears to be intermediate with Ipomoea hastigera .

Figures

Figure 3. Sepals of Ipomoea species. A I. pauciflora B I. bernoulliana C I. tentaculifera D I. hartwegii E I. murucoides F I. pantanalensis G I. hederacea H I. funis. Photographs of A (Harling et al. 15403) B (Standley 27496) C (Pringle 6702) D (Santos Martinez 2228 E (Pringle 6066) F (Pott 6399) G (McCarthy s. n.) H Andrieux 600 by John Baker.