Hydraena luridipennis MacLeay, 1873

PERKINS, PHILIP D., 2007, A revision of the Australian species of the water beetle genus Hydraena Kugelann (Coleoptera: Hydraenidae), Zootaxa 1489 (1), pp. 1-207 : 54-56

Hydraena luridipennis MacLeay, 1873

(Figs. 86, 91, 92, 214, 238)

Hydraena luridipennis MacLeay, 1873: 133 (Lectotype designated by Zwick, 1977).

Hydraena acutipennis Fairmaire, 1879: 81 . Syn. Zwick, 1977: 151.

Hydraena torrensi Blackburn, 1888: 836 . Syn. Orchymont, 1938: 426; Zwick, 1977: 151.

Hydraena selecta Deane, 1933: 27 . Syn. Zwick, 1977: 151.

Hydraena plenipennis Deane, 1933: 27 . Syn. Orchymont, 1938: 426; Zwick, 1977: 151.

Hydraena luridipennis MacLeay ; Zwick, 1977: 151.

Type Material. Lectotype (male): Queensland, Gayndah [ca. 25° 37' S, 151° 37' E] [labels: Hydraena luridipennis MacLeay Lectotype P. Zwick 1975 [written on card with specimen] // Gayndah // LECTOTYPE designated by // On permanent loan from MACLEAY MUSEUM University of Sydney // Aedeagus drawn by P. D. Perkins // DIGITAL IMAGE captured 2004 P. D. Perkins // Hydraena luridipennis MacL . Lecto-Typus des. P. Zwick 1977] [three paralectotypes and labels on same pin]. Deposited in the ANIC.

Differential Diagnosis. Recognized by the combination of the large body size (ca. 2.25 mm), the shape and relative size of the pronotum, the very small plaques, and the unique shapes of the male metatibiae and the female elytral apices (Fig. 86). The metatibiae of males are weakly arcuate on the lateral surface, the inner surface is gently sinuate, and bears a moderately sparse brush of hairs over the basal 2/3. Females are immediately recognized by the uniquely shaped, triangularly produced elytral apices. The body size, dorsal sculpture and plaques are similar to H. simplipes and H. impercepta , but males of those species have differently shaped metatibiae, and females have non-modified elytral apices. The aedeagi of the three species are quite distinct (Figs. 91–94, 96).

Description. Size: Lectotype (length/width, mm): body (length to elytral apices) 2.25/0.91; head 0.37/ 0.49; pronotum 0.53/0.68, PA 0.55, PB 0.63; elytra 1.42/0.91. Dorsum brown, except head and pronotal disc dark brown or piceous, legs and palpi brown, distal 1/2 of last palpomere darker.

Frons punctures ca. 1xef; interstices ca. 0.5 to 1xpd. Clypeus microreticulate laterally, finely sparsely punctate medially. Mentum weakly shining, very finely sparsely punctate; postmentum microreticulate. Genae raised, lacking posterior ridge. Pronotum densely coarsely punctate, punctures on disc deep, ca. 3xpd those of frons, interstices varying from narrow walls to ca. 1xpd; PF1 absent; PF2 moderately deep, separated by about width of a fovea; PF3 and PF4 moderately deep, broad.

Elytral punctures slightly smaller than largest pronotal punctures. Intervals not raised, shining, width ca. 1.5xpd. Interstices between serial punctures 1xpd or smaller. Apices (male) in dorsal aspect separately rounded, in posterior aspect margins forming marked angle with one another.

Plaques indistinct (see notes). P1 laminate; median carina angulate in profile. P2 narrow, l/w ca. 4/1, sides parallel or slightly diverging toward blunt apex. No midlongitudinal carina between mesoventral intercoxal process and median triangular depression. Cuticle non-asperite under hydrofuge pubescence. AIS flat, width at arcuate posterior margin ca. 3x P2. Profemur (male) with pointed tubercle on medial surface at about basal 1/3. Protibia straight, widest subapically. Metatibia weakly arcuate on lateral surface, inner surface gently sinuate, with brush of hairs over basal 1/2. Abdominal apex markedly asymmetrical.

Aedeagus (Fig. 91, 92) main-piece tubular, with strong, arrowhead-shaped process on right side of apical part; left paramere slightly cupped, with four groups of setae; right paramere lobe-shaped apically, with two groups of setae. Female last tergite with apicomedian incision; submarginal setae stout, forming a row, tips blunt and angled ventrad; gonocoxite as illustrated (Fig. 214).

Distribution. Widely distributed in the eastern one-half of Australia, with very disjunct locality records in western Western Australia and northern Northern Territory (Fig. 238).

Remarks. The metaventral depression is wider and deeper in males than in females. In both sexes the plaques are reduced to a very small oval or circular area on each side of the depression, near the base; in some specimens the plaques may be difficult to see, but are indicated by the less dense hydrofuge hairs. Females have simple legs, but are immediately recognized from all others by the triangularly produced elytral apices (Fig. 86). The two very disjunct localities, Badgingarra (Western Australia) and Humpty Doo (Northern Territory) are represented by a single male each. The synonyms of this species are thoroughly discussed by Zwick (1977).

Material Examined (1,670): Australian Capital Territory, 15.5 km W Canberra, Murrumbidge River, elev. 579.12 m, 35° 18' S, 149° 8' E, 2 January 2001, W. D. Shepard (8 MCZ); 2.5 km S of Tharwa, Naas River, 35° 31' S, 149° 4' E, 31 December 2000, W. D. Shepard (8 MCZ); 30 mi. SW Canberra, 36° 2' S, 149° 10' E, 1–30 January 1961, C. Watts (95 SAMA); 5 km SW Boorowa, Boorowa River, 34° 22' S, 148° 47' E, 30 December 2000, W. D. Shepard (88 MCZ); 7 km NW Tharwa, Paddy's River, 35° 31' S, 149° 4' E, 1 January 2001, W. D. Shepard (28 MCZ); Bendora Dam, 35° 27' S, 148° 50' E, 3 February 1966, Britton & Musik (1 ANIC); Black Mtn., light trap, 35° 16' S, 149° 6' E, 13 March 1968, M. S. Upton (1 ANIC); same locality, 35° 16' S, 149° 6' E, 14 March 1968, M. S. Upton (1 ANIC); same locality, 35° 16' S, 149° 6' E, 26 March 1968, M. S. Upton (1 ANIC); same locality, 35° 16' S, 149° 6' E, 29 March 1968, M. S. Upton (3 ANIC); Blundells Ck. 3 km E of Piccadilly Circus, flight intercept window/trough trap, elev. 850 m, 35° 22' S, 148° 50' E, 1 January–30 March 1984, Weir, Lawrence, Johnson (22 ANIC); Canberra, 35° 22' S, 149° 5' E, 10–11 March 1956, E. S. Brown (8 BMNH); same locality, 35° 22' S, 149° 5' E, 1–30 January 1961, C. Watts (2 SAMA); Canberra (Farrer), 35° 22' S, 149° 5' E, 30 December 1980, D. C. F. Rentz (1 ANIC); Canberrra, stream entering L. Burley-Griffin, 35° 18' S, 149° 7' E, 30 March 1965, M. E. Bacchus (MEB 221) (16 BMNH); Cotter River, 35° 30' S, 148° 50' E, 20 December 1965, E. Britton (1 ANIC); Gudgenby River, at Glendale, quartz gravel at edge of small river in pasture, elev. 875 m, 35° 41' S, 149° 0' E, 5 April 2001, N. Porch (29) (38 ANIC); Kambah Pool, pool, 35° 24' S, 149° 1' E, 26 November 1998, C. Watts (4 SAMA); nr. Canberra, 35° 22' S, 149° 5' E, 28 March 1965, M. E. Bacchus (MEB 219) (10 BMNH); Orroral R., 20 km S Tharwa, 35° 39' S, 148° 59' E, 3 January 2001, A. Glaister (3 SAMA); Paddy River, 8 km NW Tharwa, 35° 27' S, 149° 1' E, 1 January 2001, A. Glaister (1 SAMA); Paddy's River, nr. Murray's Corner, 35° 25' S, 148° 57' E, 3 April 1965, M. E. Bacchus (MEB 232) (1 BMNH); same locality, 35° 25' S, 148° 57' E, 3 April 1965, M. E. Bacchus (MEB 231) (12 BMNH); Piccadilly Circus, flight intercept window/trough trap, elev. 1240 m, 35° 22' S, 148° 48' E, 1 January–30 March 1984, J. Lawrence, T. Weir, M-L. Johnson (58 ANIC); Sullivans Creek, ANU Campus, Berlesate ANIC 1093, Azolla on pond, 35° 17' S, 149° 7' E, 18 November 1989 (3 ANIC); Tharwa, Tharwa Bridge Reserve, Murrumbidge River, 35° 31' S, 149° 4' E, 31 December 2000, W. D. Shepard (6 MCZ); Turner, 35° 16' S, 149° 7' E, 20 November 1966, K. Pullen (2 ANIC); Waramanga, 35° 21' S, 149° 4' E, 1–30 April 1973, T. Bellas (1 ANIC); Weston, 35° 20' S, 149° 3' E, 27 December 1971, T. Bellas (1 ANIC); Wombat Creek, 6 km NE of Piccadilly Circus, flight intercept window/trough trap, elev. 750 m, 35° 19' S, 148° 51' E, 1–30 March 1985, Weir, Lawrence, Johnson (14 ANIC); New South Wales, 1.6 km W Laurieville, Jerrabatgulla Creek, elev. 652.5768 m, 35° 38' S, 149° 36' E, 4 January 2001, W. D. Shepard (31 MCZ); 102 km N from Broken Hill, Fowler’s Gap, at light, 31° 5' S, 141° 42' E, 15 January 1981, Hangay & Vojnits (16 MHNG); 113 km W by S Cobar, coll. at light, 31° 35' S, 144° 39' E, 28 November 1981, J. C. Cardale (1 ANIC); 14 km W Delagate, 37° 2' S, 148° 44' E, 4 November 1997, C. Watts (1 SAMA); 34 km SE Braidwood, hwy. 52, Cabbage Tree Creek, wet sclerophyll forest, among algae, stones and leaves along margin of shaded stream, 35° 35' S, 150° 1' E, 2 April 1996, R. E. Roughley (1 NMW); 3 km SW of Wingello, ex. pool, 33° 24' S, 151° 22' E, 20 February 1974, S. Misko (2 ANIC); Armidale, 30° 30' S, 151° 39' E, 21 March 1963, C. Watts (1 SAMA); Barrington, 31° 58' S, 151° 54' E, 17 August 1997, C. Watts (3 SAMA); Barrington Tops, elev. 1524 m, 31° 59' S, 151° 27' E, 1–28 February 1932, P. J. Darlington (1 MCZ); Beaury Creek, adjacent Woodenbong State Forest, ca. 8 km SW Woodenbong, ca. 120 km SW Brisbane, Residual pools and small rock pools in river bed (Beaury Creek), in cultivated cattle country, 28° 24' S, 152° 32' E, 25 August 2004, M. A. Jäch (AU 14) (10 NMW); Blue Mountains, 33° 39' S, 150° 17' E, 1–30 January 1932, P. J. Darlington (1 MCZ); Blue Mountains, Foulcon Bridge, elev. 500 m, 33° 36' S, 150° 25' E, 15 January 1993, Wewalka (1) (1 NMW); Blue Mountains National Park, Perry’s Lookdown campground, ca. 8 km NE Blackheath, UV light, elev. 900 m, 33° 37' S, 150° 17' E, 15–16 December 1990, D. A. Pollock & L. A. Reichert (1 NMW); Boolijah Creek, 8 km E Sassafras, 35° 36' S, 150° 18' E, 24 November 2001, A. Glaister (1 SAMA); Broken Hill [no date], 31° 58' S, 141° 27' E ,, R. J. Burton (1 SAMA); Bulla Bulla tank, 31° 33' S, 147° 44' E, 19 March 1963, C. Watts (2 SAMA); Collector, 34° 51' S, 149° 26' E, 1–28 February 1961, C. Watts (2 SAMA); Colo Vale, 34° 23' S, 150° 29' E, 17 January 1957, W. W. Wirth (4 USNM); Currowan Creek, 6 km N of Nelligen, 35° 34' S, 150° 9' E, 26 November 2001, A. Glaister (1 SAMA); Darling River, 29° 57' S, 146° 18' E, 1 June 1923, H. J. Carter (1 MVMA); “Darling River” [labels read Darling River, but based on geocodes this would be the Diamantinna River], shore washing [river], 23° 11' S, 141° 42' E, 1 October 1982, Endrödy-Younga (AU 24) (28 TMSA); Forbes, 33° 22' S, 148° 0' E, 15 March 1963, C. Watts (8 SAMA); Forest Reefs [no date], 33° 26' S, 149° 5' E ,, Lea (5 QDPC); Fowlers Gap, 30° 55' S, 141° 35' E, 14 August 1983, G. Hangay (12 TMSA); Fowlers Gap Res. Stn., light/water trap, 31° 5' S, 141° 42' E, 29 November–2 December 1981, I. Naumann & J. Cardale (156 ANIC); Fowlers Gap, shore washing [river], 30° 55' S, 141° 36' E, 3 October 1982, Endrödy-Younga (AU 25) (3 TMSA); Galston, 33° 39' S, 151° 4' E, 1 February 1929, H. J. Carter (1 BMNH); Gilgandra, 31° 42' S, 148° 40' E, 19 March 1963, C. Watts (1 SAMA); same locality, 31° 42' S, 148° 40' E, 19 November 1992, C. Watts (14 SAMA); Goodradigbee R. nr. Brindabella, 35° 14' S, 148° 44' E, 11 December 1973, Sue Britton (15 ANIC); Grenfell [no date], 33° 47' S, 148° 22' E ,, E. W. Ferguson Collection (2 ANIC); Guy Fawkes River, Ebor, 30° 24' N, 152° 20' E, 31 August 2004, A. Glaister (2 NMW); Hartley Vale, Blue Mts., elev. 762 m, 33° 31' S, 150° 14' E, 30 January 1932, P. J. Darlington (16 MCZ); Kootingal (6 km W), 31° 3' S, 151° 3' E, 23 February 1992, G. L. Challet (1 USNM); Kosciusko N. P., Betts Creek, alpine meadow, ex Sphagnum etc., at stream edge and in bog, FMHD #87-267, elev. 1740 m, 36° 25' S, 148° 22' E, 14 February 1987, A. Newton & M. Thayer (5 FMNH); Macquarie Pass 7 km ENE Robertson, 34° 34' S, 150° 40' E, 8 February 1984, I. D. Naumann (1 ANIC); Megalong Vy., Blue Mts., elev. 304.8 m, 33° 43' S, 150° 15' E, 20 January 1932, P. J. Darlington (36 ANIC); Menindee Lakes Pk. Trust, Caravan Pk., at light, 32° 21' S, 142° 24' E, 26 December 1973, G. F. Gross (2 SAMA); Minnamurra River, a few km N Jamberoo, 34° 38' S, 150° 46' E, 7 March 1992, H. & P. Zwick (2 PZC); Mittagong [no date], 34° 28' S, 150° 25' E ,, Lea (1 SAMA); Mogo Creek, St. Albans, site 2, 33° 12' S, 151° 5' E, 20 November 1996, J. & R. Miller (8 JMC); Mt. Kosciusko, elev. 1524–2133 m, 36° 2' S, 148° 25' E, 1–30 December 1931, P. J. Darlington (1 MCZ); Mudgee, 32° 34' S, 149° 35' E, 1 February 1935, H. J. Carter (1 UQIC); Mulwala [no date, no collr.], 35° 58' S, 146° 0' E, (1 SAMA); Muswellbrook [no date], 32° 15' S, 150° 53' E ,, E. W. Ferguson Collection (1 ANIC); Narromine [no date], 32° 13' S, 148° 14' E ,, E. W. Ferguson Collection (5 ANIC); Nepean River, 33° 43' S, 150° 39' E, 7 September 1929, H. J. Carter & F. E. Wilson (1 MVMA); Numeralla River at Numeralla, waters edge of gravel bar at margin of flowing river, elev. 730 m, 36° 10' S, 149° 20' E, 9 April 2000, N. Porch (NP 26) (2 ANIC); Nyngan, 31° 33' S, 147° 12' E, 1 December 1953 – 30 January 1954, J. W. Armstrong (15 CAS); same locality, 31° 33' S, 147° 12' E, 16 March 1963, C. Watts (3 SAMA); On bank of Darling R., 20 mi. SSW of Bourka, at light, 30° 21' S, 145° 48' E, 25 December 1973, G. F. Gross (1 SAMA); Perisher, Porcupine, in sphagnum, elev. 1800 m, 36° 24' S, 148° 24' E, 15 January 1989, P. S. Cranston (2 ANIC); Pool near Macquarie Rivulet, near Robertson, SW Wollongong, 34° 35' S, 150° 35' E, 8 March 1992, H. & P. Zwick (1 PZC); Prospect/St. Mary's Road, 33° 47' S, 150° 55' E, 24 March 1965, M. E. Bacchus (MEB 212) (1 BMNH); Rope’s Creek [no date], 33° 49' S, 150° 48' E ,, E. W. Ferguson Collection (3 ANIC); Round Hills, near Lake Cargillego, at light, 31° 41' S, 144° 18' E, 12 January 1981, Hangay & Vojnits (6 MHNG); Sawpit Ck., Kosciusko N. P., elev. 1200 m, 31° 47' S, 150° 41' E, 14 March 1979, T. A. Weir (1 ANIC); Shaw Creek, Springwood/Richmond Rd., elev. 8 m, 33° 38' S, 150° 38' E, 24 March 1965, M. E. Bacchus (MEB 214) (21 BMNH); Siding Spring Mtn., Warrumbungle N. P., Euc. forest at light, 31° 19' S, 149° 0' E, 23 November 1985, C. Reid (3 ANIC); Sydney, elev. 100 m, 33° 52' S, 151° 12' E, 11 July 1943, H. P. Chandler (3 CAS); Sydney, Griffith Collection [no date], 33° 52' S, 151° 12' E, (3 SAMA); Tenterfield, Bluff River, 28° 49' S, 151° 56' E, 29 August 2004, A. Glaister (2 NMW); Tenterfield, Dumaresq River, 28° 59' S, 151° 31' E, 28 August 2004, A. Glaister (2 NMW); The Lake Ck., 12 km NNE of Jindabyne, elev. 1160 m, 36° 25' S, 148° 38' E, 16 March 1979, T. A. Weir (1 ANIC); Tooloom Scrub via Urbenville, pyrethrum on logs, 28° 34' S, 152° 29' E, 28 September 1989, G. B. Monteith (1 QMBA); Upper Williams River, 32° 21' S, 151° 45' E, 1–30 October 1926, Lea & Wilson (2 SAMA); Valley near Blackheath, Blue Mountains, elev. 914.4 m, 33° 37' S, 150° 17' E, 23 January 1932, P. J. Darlington (1 MCZ); Wilcannia, 31° 34' S, 143° 22' E, 1–30 March 1960, C. Watts (2 SAMA); Willandra Bridge, 11 km N of Mossgriel, dry swamp, at light, 33° 16' S, 144° 34' E, 21 December 1970, Britton, Misko, Pullen (1 ANIC); Wollindibby Ck., 10 km WbyS of Jindabyne, elev. 1060 m, 36° 28' S, 148° 29' E, 15 March 1979, T. A. Weir (1 ANIC); Woodlands Creek at Kuradji Sandon Point, 34° 19' S, 150° 55' E, 2 April 2001, J. & R. Miller (1 JMC); Young (40 km N), pond, 33° 39' S, 150° 18' E, 20 February 1992, G. L. Challet (1 USNM); Northern Territory, 80 mi. NW Alice Springs, 23° 23' S, 133° 3' E, 1– 28 February 1968, C. Watts (1 SAMA); Alice Springs [no date, no collr.], 23° 42' S, 133° 52' E ,, unknown (2 ANIC); Border Waterhole, Musselbrook Ck. 15 km W by S of Musselbrook Mining Camp ( QLD), large permanent pool in gorge, muddy edges, rocky, algal growth, 18° 37' S, 137° 59' E, 14 May 1995, T. Weir (2 ANIC); Finke Gorge N. P., Kanara Ck. at The Oasis, permanent rock pools, palms, grass at edges, algal growth, 24° 4' S, 132° 44' E, 12 March 1995, T. Weir (1 ANIC); Humpty Doo (6 km E), 12° 37' S, 131° 15' E, 9 February–4 March 1987, R. I. Storey (1 QPIM); Palm Valley, Finke Gorge National Park, 24° 4' S, 132° 44' E, 10–12 August 1974, T. Weir & T. Angeles (6 ANIC); Renner Springs, 18° 19' S, 133° 48' E, 4 May 1962, C. H. S. Watts (2 SAMA); Queensland, 15 km ESE Warwick, Queen Mary Falls N. P., E Killarney, elev. 300 m, 28° 20' S, 152° 22' E, 26 August 2004, M. A. Jäch (AU 15) (1 NMW); 18 km N Mareeba, intermittent stream, 16° 58' S, 145° 25' E, 20 September 1990, D. Larson (1 MCZ); 30 km NE Brisbane, Mt. Nebo Road, Pond, ca. 200 m, Sclerophyll forest, elev. 540 m, 27° 24' S, 152° 48' E, 21 August 2004, M. A. Jäch (AU 1) (3 NMW); 30 km NW Brisbane, South Pine River, elev. 120 m, 27° 21' S, 152° 47' E, 21 August 2004, M. A. Jäch (AU 2) (7 NMW); 31 km NW by N of Longreach, 23° 13' S, 144° 4' E, 10 May 1973, M. S. Upton (1 ANIC); 4 km N Conondale, Sandy Creek, 26° 42' S, 152° 41' E, 17 August 2004, A. Glaister (3 NMW); 40 km S Beaudesert, Running Creek, along road between Mt. Chinghee N. P. and Border Ranges N. P., at 7th Bridge, stream, ca. 5–8 m wide, including rockpools, flowing through open forest ( Casuarina , Eucalyptus ), 28° 19' S, 152° 55' E, 25 August 2004, M. A. Jäch (AU 13) (9 NMW); 6 km W Dimbulah, 17° 8' S, 145° 6' E, 10 November 1990, D. Larson (1 MCZ); 60 km S Brisbane, 10 km NNW Canungra, 27° 57' S, 153° 8' E, 25 August 2004, M. A. Jäch (AU 11) (5 NMW); 8 km W Rathdowney, Logan River, 28° 12' S, 152° 47' E, 27 August 2004, A. Glaister (2 NMW); 90 km SW Brisbane, 40 km NE Warwick, 28° 3' S, 152° 22' E, 26 August 2004, M. A. Jäch (AU 17) (10 NMW); Albert River, S of Wolfdene, 30 km SE Brisbane, River (Albert River), ca. 20 m wide, slightly polluted, flowing through cultivated land, elev. 30 m, 27° 46' S, 153° 11' E, 21 August 2004, M. A. Jäch (AU 3) (2 NMW); “Big Spring”, 3 km SE Edgbaston H. S., 31 km NE Aramac, spring, 22° 45' S, 145° 25' E, 4 May 1988, W. Zeidler (5 SAMA); Biloela, 24° 24' S, 150° 30' E, 7 September 1972, H. E. Hinton (14 BMNH); Birdsville, 25° 53' S, 139° 21' E, 14 September 1968, C. Watts (2 SAMA); Bluewater, 19° 10' S, 146° 24' E, 31 October 1995, C. Watts (1 SAMA); Boggom.30 via Taroom, baited flight intercept, 057, 25° 27' S, 150° 2' E, 14 November 1996 – 30 January 1997, Cook & Monteith (4 QMBA); Boggom.8 via Taroom, baited flight intercept, 035, 25° 27' S, 150° 2' E, 12 November 1996 – 30 January 1997, Cook & Monteith (1 QMBA); Boggomoss No. 19, via Taroom, night hand collection, 034, 25° 25' S, 150° 1' E, 11 November 1996, Qld. Mus. survey (1 QMBA); Boggomoss No. 21, via Taroom, aquatic collection, 073, 25° 27' S, 150° 3' E, 11 November 1996, Thompson & Monteith (3 QMBA); Boggomoss No. 21, via Taroom, at light, 028, 25° 27' S, 150° 3' E, 11 November 1996, G. B. Monteith (1 QMBA); Borumba Dam, Yabba Creek Crossing No. 6, elev. 150 m, 26° 25' S, 152° 45' E, 18 November 1996, Endrich (26) (79 NMW); Brisbane, 27° 22' S, 152° 46' E, 23 September 1892, C. Wild (1 SAMA); same locality, 27° 22' S, 152° 46' E, 12 April 1957, Haseler (1 UQIC); same locality, 27° 22' S, 152° 46' E, 2 June 1957, Haseler (4 UQIC); Brisbane State Forest, Manorina N. P., elev. 530 m, 27° 22' S, 152° 46' E, 19 November 1996, L. Hendrich (28) (4 NMW); Bruce Hwy. nr Childers, 10 km N Howard, creeks, 25° 14' S, 152° 16' E, 17 November 1996, L. Hendrich (20 NMW); Bullaroo Creek, 150 km N of Roma, on road to Carnarvon National Park, 25° 11' S, 148° 34' E, 17 March 1992, H. & P. Zwick (1 PZC); Bunya Mt., logs and bark, 26° 52' S, 151° 40' E, 11 December 1982, Endrödy-Younga (AU 101) (1 TMSA); Bunya Mts. N. P., rainforest litter, SBP 108, elev. 1000 m, 26° 51' S, 151° 34' E, 18 August 1982, S. Peck (1 ANIC); Burnett River, Mt. Perry Rd., 25° 23' S, 151° 46' E, 15 August 2004, A. Glaister (1 NMW); C. QLD: Peawaddy Gorge, Mt. Moffatt N. P., elev. 1100 m, 24° 55' S, 148° 4' E, 26 February 1996, G. B. Monteith (1 QMBA); Camooweal, 19° 55' S, 138° 7' E, 30 April 1963, C. Watts (2 SAMA); Canungra Ck., Sarabah Bridge, 10 km SW Canungra, 28° 4' S, 153° 6' E, 22 August 2004, A. Glaister (1 NMW); Canungra Creek, 60 km SSE Brisbane, River (Canungra Creek), ca. 8–10 m wide, through cultivated land, elev. 160 m, 28° 4' S, 153° 6' E, 22 August 2004, M. A. Jäch (AU 6) (7 NMW); Carnarvon, 24° 56' S, 148° 39' E, 29 May 1954, F. A. Perkins (1 UQIC); Carnarvon Ck. 17 km E Park Office, 25° 1' S, 148° 18' E, 14 August 2004, A. Glaister (5 NMW); Carnarvon Gorge, 25° 4' S, 148° 15' E, 24 May–2 June 1954, F. A. Perkins (1 UQIC); Clermont, 22° 44' S, 147° 26' E, 1–30 December 1929, K. K. Spence (15 AMSA); Cloncurry, 20° 42' S, 140° 30' E, 29 April 1963, C. Watts (2 SAMA); Condamine R. Killarney, 27° 3' S, 149° 38' E, 6 November 1932, H. Hacker (7 QMBA); Connor’s River, 91.5 miles S Mackay, 22° 39' S, 149° 4' E, 3 September 1972, H. E. Hinton (18 BMNH); Dargonelly Rockhole, 25° 2' S, 147° 54' E, 27 February 1996, G. Monteith & C. Burwell (5 QMBA); Darr River, 31 km NW by N Longreach, 23° 40' S, 143° 50' E, 7 April 1976, D. H. Colless (1 ANIC); Edungalba, ex flight intercept trap, 23° 43' S, 149° 50' E, 1 January 1987, H. & A. Howden (1 ANIC); Edungalba, 23° 43' S, 149° 50' E, 14 December 1988, H. & A. Howden (1 CMNO); Eumamurrin Creek, 53 km N of Roma, on road to Carnarvon National Park, 26° 14' S, 148° 42' E, 17 March 1992, H. & P. Zwick (34 PZC); Finch Hatton, Palmtree Creek, 21° 6' S, 148° 37' E, 5 September 1972, H. E. Hinton (1 BMNH); Gatton, 27° 32' S, 152° 3' E, 9 November 1932, [collector not given] (2 QDPC); Gayndah, 25° 37' S, 151° 37' E (1 ANIC); Girraween N. P., Bald Rock Creek at "Under-ground Creek", ex. sand/gravel streambank, 28° 51' S, 151° 57' E, 22 January 1998, P. D. Perkins (5 MCZ); Girraween N. P., Bald Rock Creek near Bald Rock camp ground, 28° 49' S, 151° 55' E, 29 August 2004, A. Glaister (2 NMW); Glengyle, 24° 47' S, 139° 35' E, 15 September 1968, C. Watts (2 SAMA); Greenvale, 150 km NW Charters Towers, elev. 500 m, 19° 0' S, 144° 58' E, 19 January 1993, Wewalka (1 NMW); Greenvale 70 km SW, at light, 19° 23' S, 144° 37' E, 10–20 October 1995, A. J. Watts (1 SAMA); Indooroopilly, pan traps along creek, 27° 30' S, 152° 59' E, 28 February–9 March 1984, L. Masner (2 CMNO); Julia Creek, 20° 3' S, 141° 12' E, 24 April 1963, C. Watts (15 SAMA); Kenilworth, Obi Obi Creek, elev. 150–200 m, 26° 36' S, 152° 44' E, 18 November 1996, L. Hendrich (27) (6 NMW); Lamington N. P., Comera River, 80 km S Brisbane, 28° 8' S, 153° 9' E, 24 August 2004, M. A. Jäch (AU 10) (10 NMW); Lamington N. P., Moran’s Creek, 80 km S Brisbane, elev. 760–820 m, 28° 14' S, 153° 8' E, 24 August 2004, M. A. Jäch (AU 9) (2 NMW); MEQ: Boomer Range, Site 8, open forest, intercept trap, elev. 220 m, 23° 12' S, 149° 46' E, 16 December 1999 – 22 March 2000, G. B. Monteith (1 QMBA); MEQ: Mt. Bluffkin, 7 km NNE, on road Brigalow, intercept trap, 22° 36' S, 149° 14' E, 22 March–2 June 2000, Monteith & Cook (1 QMBA); Mia Mia State Forest, between Finch Hatton & Pinnacle, Cattle Creek, 21° 14' S, 148° 47' E, 16 March 1997, P. Zwick (25 PZC); Nanango, Barkers Creek [no date], 26° 40' S, 151° 59' E ,, L. Wassell (1 UQIC); Noccundra, at light, 27° 49' S, 142° 35' E, 2 May 1968, R. Lewis (1 ANIC); Pioneer R. nr. Mirani, 21° 9' S, 149° 13' E, 4 September 1972, H. E. Hinton (2 BMNH); Queen Mary Falls N. P., E Killarney, 15 km ESE Warwick, elev. 770 m, 28° 20' S, 152° 22' E, 26 August 2004, M. A. Jäch (AU 16) (2 NMW); Reid R. E of Charters Towers, 19° 45' S, 146° 48' E, 4 May 1998, C. H. S. Watts (1 SAMA); Richmond, 20° 56' S, 143° 20' E, 1 December 1917, [collector unknown] (2 NMW); same locality, 20° 56' S, 143° 20' E, 20 February 1918, [collector unknown] (7 NMW); same locality, 20° 56' S, 143° 20' E, 23 February 1918, [collector unknown] (1 NMW); same locality, 20° 56' S, 143° 20' E, 12 March 1918, [collector unknown] (2 NMW); same locality, 20° 56' S, 143° 20' E, 20 May 1918, [collector unknown] (4 NMW); Rockhampton, Bruce Hwy. near Raglan, 12 Mile Creek, elev. 20 m, 23° 22' S, 150° 30' E, 14 November 1996, L. Hendrich (24) (31 NMW); Running Creek Road, SE Rathdowney, ca. 25 km SSE Beaudesert, small road side ditch (remnants of small swampy meadow), 28° 15' S, 152° 53' E, 25 August 2004, M. A. Jäch (AU 12) (3 NMW); South of Charleville, 26° 24' S, 146° 20' E, 9 May 1973, M. S. Upton (1 ANIC); Stanthorpe, 28° 39' S, 151° 56' E, 1–30 January 1961, C. Watts (5 SAMA); Tambourine Mt., 27° 53' S, 153° 11' E, 20 January 1987, W. Dressler (1 ANIC); Taroom Dist. Boggom.11, 162, 25° 28' S, 150° 3' E, 14 June 1996, R. Kristensen (14 QMBA); Taroom Dist. Boggom.3, 163, 25° 26' S, 150° 1' E, 14 June 1996, R. Kristensen (11 QMBA); Taroom District, Boggomoss 21, open forest, sieved leaf litter, QM berlesate no. 912, elev. 200 m, 25° 27' S, 150° 3' E, 10 November 1996, G. B. Monteith (1 QMBA); Townsville, 19° 15' S, 146° 48' E, 24 March 1932, P. J. Darlington (22 MCZ); same locality, 19° 15' S, 146° 48' E, 16–22 March 1965, J. Balogh (1 HNHM); Townsville [no date], 19° 15' S, 146° 48' E ,, E. W. Ferguson Collection (1 ANIC); same locality, 19° 15' S, 146° 48' E, 19 April 1963, C. Watts (6 SAMA); Upper Brookfield, 27° 28' S, 152° 51' E, 10 December 1962, G. Monteith (1 UQIC); Young’s Crossing, Petrie, 27° 16' S, 152° 57' E, 2 October 1959, I. C. Yeo (2 UQIC); South Australia, 3rd Spr. Willigon Ck, Flinders Rg., creek spring, 31° 0' S, 138° 38' E, 15 December 1976, J. E. Bishop & J. H. Diener (1 SAMA); Adelaide, 34° 55' S, 138° 36' E, 1– 30 March 1956, C. Watts (3 SAMA); Adelaide [no date], 34° 55' S, 138° 36' E ,, Blackwell (1 BMNH); Alligator Gorge, 32° 44' S, 138° 4' E, 1–30 June 1959, C. Watts (1 SAMA); Balcanoona Ck., Nth. Flinders, 30° 33' S, 139° 20' E, 24 July 1976, P. Suter, B. Mitchell, R. Marchant (1 SAMA); Chambers Gorge, 30° 58' S, 139° 16' E, 1–30 September 1983, C. Watts (2 SAMA); Cooper Creek, Embarka W. H., 27° 41' S, 140° 12' E, 21 July 1994, Aust. water quality centre 3784 (1 SAMA); Enorama Ck. on Aroona Valley Rd., Nth. Flinders, 31° 19' S, 138° 41' E, 26 July 1976, P. Suter, B. Mitchell, R. Marchant (1 SAMA); Lake Giles Tank, Gawler Ranges, Eyre Pen., 32° 38' S, 136° 53' E, 12 December 1976, J. E. Bishop & J. H. Diener (2 SAMA); Lake Pinpa, 40 km E Frome Downs Station, on lake surface, 31° 13' S, 140° 11' E, 1–30 August 1976, N. Pledge (1 SAMA); Mt. Compass, 35° 19' S, 138° 36' E, 1–30 December 1960, C. Watts (2 SAMA); Mt. Gambier, 37° 50' S, 140° 45' E, 1–30 December 1961, C. Watts (5 SAMA); Mt. Lofty Rgs. [no date], 34° 58' S, 138° 42' E ,, A. H. Elston (10 AMSA); Mybonga [no date],, R. F. Kemp (2 AMSA); Old Wirrialpa Spr., Flinders Rg., 31° 4' S, 138° 54' E, 16 December 1976, J. E. Bishop & J. H. Diener (7 SAMA); Orlunga Ck., Nt. Flinders, 31° 5' S, 138° 33' E, 25 July 1976, P. Suter, B. Mitchell, R. Marchant (1 SAMA); Parachilna Ck., nr. Mt. Mary, Nth. Flinders, 31° 8' S, 138° 31' E, 26 July 1976, P. Suter, B. Mitchell, R. Marchant (1 SAMA); Quorn [no date], 32° 20' S, 138° 2' E ,, A. H. Elston (1 MVMA, 2 SAMA); Scotts Creek Park, near Adelaide, 35° 5' S, 138° 41' E, 12 December 2001, M. Balke (5 NMW); Warburton R., 2 km S White Bull Yard, Kalamurina Stn., 27° 54' S, 137° 54' E, 7 October 1999, WHC exped. (1 SAMA); Williamstown, 34° 40' S, 138° 53' E, 1–30 September 1961, C. Watts (1 SAMA); same locality, 34° 40' S, 138° 53' E, 1–30 October 1961, C. Watts (1 SAMA); Victoria, 10 km NE Mirranatwa, small creek bed/drying pool, stony, dead leaves, dry sclerophyll forest, 37° 36' S, 142° 13' E, 12 October 1997, C. Watts (1 SAMA); 20 km NNE of Licola, Bennison Plains, 37° 28' S, 146° 43' E, 30 November 1976, A. A. Calder (12 MVMA); 21 miles E of Echuca, in irrigation channel, 36° 8' S, 144° 45' E, 27 October 1967, Britton & Misko (1 ANIC); 4 km W of Violet Town, ex small pool, 36° 37' S, 145° 44' E, 22 November 1973, S. Misko (8 ANIC); 8 km SW Moyston, 37° 17' S, 142° 39' E, 16 November 1969, E. F. Riek (5 ANIC); Acheron River, Narbethong, 37° 33' S, 145° 39' E, 2 December 1998, C. Watts (1 SAMA); Alps [no date], 37° 30' S, 145° 53' E ,, C. French (1 MVMA); Avoca River at Avoca, litter and gravelly sand at edge of small river; pasture, elev. 225 m, 37° 5' S, 143° 28' E, 12 March 2000, N. Porch (71) (1 ANIC); Bacchus Marsh, 12 km W by N of, shallow, grassy, roadside ditch, and from litter and mud at edge of ditch, elev. 455 m, 37° 38' S, 144° 17' E, 17 September 2000, N. Porch (4) (16 ANIC); Bannimboola Creek, 10 km WNW Dartmouth, 36° 30' S, 147° 30' E, 29 October 1976, A. A. Calder (2 MVMA); Black Calf Creek near Mitton’s Bridge, 13 km south of Kinglake, 37° 35' S, 145° 18' E, 29 September 1972, P. Zwick (1 PZC); Boggy Creek at Princes Hwy. bridge, Nowa Nowa, cobbles and muddy gravel at edge of small river; dry sclerophyll forest, pasture, elev. 5 m, 37° 43' S, 148° 5' E, 19 September 1999, N. Porch (59) (1 ANIC); Brisbane Ranges National Park, 8.2 km W Rowsley Trig., litter and mud at edge of small firedam in dry sclerophyll forest, elev. 405 m, 37° 44' S, 144° 16' E, 1 January 2000, N. Porch (60) (1 ANIC); Caladonia Fen, 2.5 km NE Snowy Range airstrip, elev. 1300 m, 37° 20' S, 146° 45' E, 8 December 1998, N. Porch (1 ANIC); Caledonia Fen, 3 km NNW of Snowy Range Airfield, Alpine National Park, small pool in drying fen; montane sclerophyll forest, elev. 1300 m, 37° 19' S, 146° 44' E, 22 February 2001, N. Porch (23) (8 ANIC); Dandenong Creek near mouth into sea, 37° 56' S, 145° 12' E, 1 March 1972, P. Zwick (1 PZC); Dinner Plain to Mt. Hotham, shallow macrophyte rich pond in subalpine snow gum ( E. pauciflora ) forest, elev. 1630 m, 37° 0' S, 147° 10' E, 16 December 2001, N. Porch (89) (5 ANIC); Dondangadale, 36° 47' S, 146° 40' E, 1 November 1955, Neboise (8 MVMA); East Pomborneit, 24 km ESE Camperdown, temporary pond, 38° 17' S, 143° 22' E, 1 August 1978 – 28 February 1979, P. S. Lake (6 ANIC); East Tangil River, 37° 50' S, 146° 12' E, 8 November 1997, C. Watts (1 SAMA); Eel Hole Creek at Spilway. S47 LaTrobe C. Survey, 38° 17' S, 146° 22' E, 8 December 1978, [collector not given] (1 MVMA); Fitzroy River [no date, no collr.], 38° 9' S, 141° 42' E, (4 BMNH); Fryer’s Creek near mouth into Lake Eildon, 37° 11' S, 145° 53' E, 3 October 1972, P. Zwick (1 PZC); Godfrey’s Creek above AI Mine Settlement, 38° 36' S, 143° 54' E, 11 October 1972, P. Zwick (1 PZC); Goulbourn River, at Goulbourn Valley Hwy., 8.0 km SE. Seymour, sandy mud at edge of large slow river, elev. 145 m, 37° 5' S, 145° 12' E, 11 August 1999, N. Porch (70) (1 ANIC); Grampians, 37° 15' S, 142° 26' E, 1–28 February 1963, C. Watts (1 SAMA); Halls Gap, 37° 10' S, 142° 33' E, 13 January 1997, C. Watts (1 SAMA); Healesville, Dandenong Ranges, 37° 39' S, 145° 32' E, 1–30 December 1968, C. Watts (5 SAMA); same locality, 37° 39' S, 145° 32' E, 15 January 1997, C. Watts (1 SAMA); Jamieson River at Jamieson, 38° 35' S, 143° 53' E, 2 October 1972, P. Zwick (1 PZC); Jordan River/ BB Creek Junction, Jericho, NMV Survey Dept. GRES T 10 no. 1, 37° 39' S, 146° 17' E, 28 November 1976, [collector not given] (2 MVMA); Kangaroo Creek on Springhill Rd., 5.8 km E Glenlyon, amongst weeds in shallow water at edge of creek (perennial) pools, dry sclerophyll forest, elev. 510 m, 37° 18' S, 144° 18' E, 31 January 1999, N. Porch (NP 24a) (1 ANIC); Kiewa Valley Hwy. n. Redbank, 36° 56' S, 143° 19' E, 13 February 1998, L. Hendrich (2 NMW); Latrobe River at Piedmont, gravel at edge of river at pasture/forest boundary, elev. 255 m, 37° 53' S, 145° 54' E, 1 January 2003, N. Porch & N. Krause (NP 54) (1 ANIC); Little River at Bacchus Marsh/Geelong Road, sandy gravel and muddy sand at edge of still pool of perennial creek in riparian forest remnants, elev. 105 m, 37° 50' S, 144° 23' E, 20 October 1999, N. Porch (36) (4 ANIC); Loddon Falls, base of, NE of Glenlyon, sandy gravel at edge of waterfall splashpool in scrubby gorge [stream], elev. 470 m, 37° 16' S, 144° 14' E, 16 March 2001, N. Porch & K. Sniderman (19) (1 ANIC); Loddon River at Newstead, gravel at edge of small river flowing through pasture, elev. 210 m, 37° 6' S, 144° 3' E, 27 November 1999, N. Porch (47) (2 ANIC); Macalister/Thomson R. Jnc., 37° 46' S, 146° 40' E, 19 February 1977, A. A. Calder (1 MVMA); Marysville, 37° 31' S, 145° 45' E, 1–30 December 1968, C. Watts (1 SAMA); McFarlane Saddle, 1.75 km WNW of Trapyard Hill, roadside firedam, mud and shallow water at the edge of a small clay based firedam lacking macrophytes, elev. 1495 m, 37° 28' S, 147° 50' E, 28 November 1999, N. Porch & N. Krause (43) (15 ANIC); Melbourne [no date], 37° 58' S, 144° 58' E, (1 BMNH); Merriman Creek, 2.0 km NW Seaspray, gravel at base of gravel bank of small creek; pasture, elev. 5 m, 38° 21' S, 147° 9' E, 7 April 1999, N. Porch & N. Krause (50) (3 ANIC); Mitta Mitta River, 10 km NW of Dartmouth Dam, 36° 30' S, 147° 26' E, 1 November 1976, A. A. Calder (1 MVMA); Mitta Mitta River at Mitta Mitta, 36° 32' S, 147° 22' E, 1 December 1973, Neboiss & Malcolm (1 MVMA); Mitta Mitta River/Snowy Creek Junction, 36° 30' S, 147° 26' E, 3 November 1976, A. A. Calder (1 MVMA); Mogg’s Creek, 38° 27' S, 144° 3' E, 1 January 1940, P. Radford (1 MVMA); Morses Creek, Bright Caravan Park, Bright, gravel and mud at edge of small stream in developed park, elev. 330 m, 37° 44' S, 144° 7' E, 13 December 2001, N. Porch (80) (1 ANIC); Mt. Disappointment Plateau, Mountain Road/Bruce Creek Road Junction, muddy litter at edge of firedam in dry sclerophyll forest, elev. 605 m, 37° 23' S, 145° 6' E, 17 October 1999, N. Porch (48) (2 ANIC); Nolans Creek picnic ground, 1 km SSW of Mt. Wilson, gravel and litter at edge of small forest stream flowing through sclerophyll forest, elev. 645 m, 37° 27' S, 144° 14' E, 6 January 2002, N. Porch (81) (1 ANIC); Otways, Carisbrook Creek near mouth into sea, 38° 38' S, 143° 46' E, 14 November 1972, P. Zwick (1 PZC); Ovens River at Harrietville, 36° 53' S, 147° 4' E, 23 November 1972, P. Zwick (1 PZC); Settling Ponds Dam, at Junction of Settling Ponds Track and Hudbrick Track, gravelly mud at edge of small dam in dry sclerophyll forest, elev. 260 m, 37° 7' S, 143° 40' E, 9 June 2001, N. Porch (74) (11 ANIC); Shaw Creek, Bennison Plains, NMV Survey Dept. GRES Mc9 no. 8, 37° 32' S, 146° 40' E, 30 November 1976, [collector not given] (5 MVMA); Simpson Creek, 12 km SW Orbost, 37° 44' S, 148° 20' E, 30 November 1998, C. Watts (1 SAMA); Six Mile Creek, Dartmouth Survey, Locality GA, 37° 12' S, 146° 23' E, 7 April 1973 (1 MVMA); Thompson River, Low Saddle Trk, NMV Survey Dept. GRES T 15 no. 3, 37° 41' S, 146° 13' E, 24 October 1977, [collector not given] (1 MVMA); Thomson River, Moe/ Walhalla Rd., NMV Survey Dept. GRES T 21 no. 5, 37° 41' S, 146° 13' E, 1 December 1976, [collector not given] (2 MVMA); Thomson River, Thomson Valley Rd., NMV Survey, Dept. GRES T 1, no. 2 kick, 37° 41' S, 146° 13' E, 25 November 1976, unknown (1 MVMA); Thomson River, Thomson/ Jordan Divide Rd., NMV Survey Dept. GRES T 21 no. 5, 37° 41' S, 146° 13' E, 2 December 1976, [collector not given] (1 MVMA); Toorongo River 1 km S of Toorongo Rd. S12, LaTrobe C. Survey, 37° 47' S, 146° 4' E, 29 November 1978 (2 MVMA); Wellington/Carey Junction, NMV Survey Dept. GRES Mc17 no. 1, 37° 20' S, 146° 50' E, 2 December 1976, [collector not given] (1 MVMA); Western Tyers River on Xmas Ck. Rd. S52 LaTrobe C. Survey, 37° 54' S, 146° 16' E, 3 December 1978, [collector not given] (2 MVMA); Woori Yallock Creek on road to Warburton, 37° 44' S, 145° 29' E, 4 November 1972, P. Zwick (1 PZC); Yellow Creek, at crossing of Ovens Highway, Wangaratta, coarse sand and gravel at wet margins of gravel bar of red gum lined creek in pasture, elev. 145 m, 36° 21' S, 146° 20' E, 15 December 2001, N. Porch (88) (1 ANIC); Western Australia, Badgingarra, Hill River, 30° 23' S, 115° 3' E, 10 September 1990, D. Larson (1 MCZ).