Hycleus colligatus (Redtenbacher, 1850)

Serri, Sayeh, Bologna, Marco A. & Riccieri, Alessandra, 2020, Revision of the Hycleus sexmaculatus species group (Coleoptera: Meloidae Mylabrini), Zootaxa 4790 (1), pp. 1-42 : 16-19

Hycleus colligatus (Redtenbacher, 1850)

Figs 5, 17, 55–62

Mylabris colligata Redtenbacher (in Kollar & Retenbacher), 1850: 49 .

Mylabris javeti Marseul, 1870: 88 syn. n.

Mylabris javeti umbilicatus Kaszab, 1958: 291 syn. n.

Hycleus colligatus, Bologna & Turco, 2007: 17 .

Hycleus javeti, Bologna & Turco, 2007: 17 .

Type material. 2 ♂ 1 ♀ syntypes, “ colligata // Kots./ m” ( NHMW) .

Four syntypes of Mylabris javeti (possibly other specimens close in the box in the Marseul’s collection to the holotype could be syntypes) ( MNHN); 1♂ syntype, “ Persia / settent./ 1862–63 / Col. G. Doria // Collect./ Türk // Ja- veti Mars./ type/ Persia Heyd. // Javeti” ( NHMW) .

Paratypes of Mylabris javeti umbilicata , 1 ♂ “ Nuristan (Afghan.) 1935/ Expedit. Dr. Sceibe //Pargil n./ Patschlig- il/ 12.7.35// Myl. colligata Redtb. // Deutsch. Entomol./Institut Berlin// Paratypus / Mylabris javeti / ssp. umbilicata m./ det. Dr. Kaszab, 1956” ; 1 ♂ “J. Klapperich/ Umgeb. Kabul / 1740 m, 24.6.52/ O- Afghanistan // Paratypus 1 ♂ / Mylabris javeti / ssp. umbilicata m./ det. Dr. Kaszab, 1956” ; 1 ♀ “J. Klapperich/ Bashgul Tal 1200 m / Nuristan 11.5.53/ Afghanistan // Paratypus / Mylabris javeti / ssp. umbilicata m./ det. Dr. Kaszab, 1956” ( HNHM) .

Type locality: “Südpersien“ ( Kollar & Redtenbacher 1850).

Additional material examined. Afghanistan. 5 ♂ 6 ♀ Borgh , Logar, 19.VII.1957, no. 246, leg. Lindberg ( HNHM) ; 1 ♂ Ghazni, 30 km SW Moqur , 1900 m, 10. V .1973, leg. Kabakov ( HNHM); 1♀ Ghazni, 50 km NW Moqur , 2200 m, 10. VI .1972, leg. Kabakov ( HNHM); 1 ♂ E Afghanistan, Gulbahar , 1700 m, 15. VI .1956, leg. Amsel ( HNHM); 1 ♂ Heart , 970 m, 15. V .1956, leg. H. G. Amsel ( HNHM); 1 ♀ Kouh- Djaouz , 20 km de Kaboul, 1820 m, 31. V .1960, no. 847, leg. Lindberg ( HNHM); 1 ♂ umgeb. Kabul, 1740 m, 24. VI .1952, leg. Klapperich ( HNHM); 1 ♀ Qal′eh Lakou, aff. Val. Kaboul, 7.VII.1957, no. 254, leg. Lindberg ( HNHM) ; 1 ♂ Laghman, Samakat , 900 m, 24. IV .1972, leg. Kabakov ( HNHM); 1 ♀ Nangarhar prov., (112) Darunta , 580–600 m, 3. V .1967, leg. Povolny ( HNHM); 1♀ Nuristan, Bashgaltal, 120 0m, 11. V .1953, leg. Klapperich ( HNHM); 1 ♂ Nuristan, Parigil n., Patschlig- il, 12.VII.1935, leg. Scheibe ( HNHM) ; 1 ♂ Nangarhar, Dasht-e Gamberay , 650 m, 8. V .1974, no. 82, leg. Papp ( HNHM); 2 ♂ 1♀ Nuristan, Parigil n. Patsdeligil , 12.VII.1935, Exp. Dr. Scheibe ( HNHM) ; 3♀ Darreh Gazak , 20 km S Oubeh, 1370 m, 10. VI .1962, leg. K. Lindberg ( HNHM). Caucasus. 1 ♂ R . Oberthur, Eing. Nr. 4. 1956 ( HNHM) . Iran. 1 ♂, Alborz prov., Aderan, Karaj , 23.VII.1946 ( HMIM) ; 1 ♀ Alborz prov., Eshtehard, 26. V .1968, leg. Ayatollahi ( HMIM); 1 ♂ Bushehr prov., Borazjan, Asaluyeh , 40 m, 23. IV .1977, leg. Pazuki & Hashemi ( HMIM); 1 ♂ 2 ♀ Bushehr prov., Bushehr, Ahram, Tangestan , 100 m, 4–5. V .1995, leg. Parchami, Badii & Ardeh ( HMIM); 1 ♂ Bushehr prov., Borazjan , 19.III.1953, leg. Mirsalavatian ( HMIM) ; 6 ♂ 7♀ Bushehr prov., Dylam, 1, 9. V .1969, leg. Zairi & Ayatollahi ( HMIM); 14 ♂ 2 ♀ Esfahan prov., Delijan, 2. VI .1970, leg. Mirzayans & Abai ( HMIM); 3 ♂ 1♀ Esfahan prov., Delijan , 1700 m, 8. VI .1986, leg. Mirzayans & Hashemi ( HMIM); 1 ♂ Esfahan prov., Esfahan, V .1948, leg. Makoui ( HMIM); 1 ♀ Esfahan prov., Khansar, 16. VI .1948, leg. Alexandrov ( HMIM); 1 ♂ 1 ♀ Esfahan prov., Khansar , Golestan kuh, 2700 m, 3/ 4.VII.1983, leg. Mirzayans & Broumand ( HMIM) ; 20 ♂ 18♀ Esfahan prov., Kashan- Niasar, Sericheh , 1650 m, 9–11. VI .1984, leg. Pazuki & Hashemi ( HMIM); 2 ♂ 4 ♀ Esfahan prov., Kashan, Rahagh , 2350 m, 25. VI .1981, leg. Hashemi ( HMIM); 15 ♂ 7♀ Esfahan prov., 25 km E Khomein , 1860 m, 2.VII.1983, leg. Mirzayans & Borumand ( HMIM) ; 1♂ 1♀ Esfahan prov., Zagros Mt. Shahreza, Koruyeh, 11. VI .2011, leg. Dalihod ( MABC); 2 ♂ [Fars prov.], zentral Iran. 20 km nordl. v. Abadeh, 2000m, leg. Vartaian ( HNHM) ; 1♂ Perisa, [Fars prov.], Abadeh , V–VI.1916, leg. P. Paschen, coll. P. Gough ( HNHM) ; 5 ♂ 5♀ Fars prov., Dadin, 8, 11. V .1968, leg. Mojib ( HMIM); 1 ♀ Fars prov., Dadin, 7. V .1965, leg. Mojib ( HMIM); 2 ♂ Fars prov., Dascht-e Arjan, 27. VI .1976, leg. Sbordoni & Rampini ( MABC); 1 ♀ Fars prov., Eghlid, 3. VI .1969, leg. Termeh & Izadyar ( HMIM); 3 ♂ 1♀ Fars prov., Farashband, Hosseinabad, 4. VI .1968, leg. Mirzayans ( HMIM); 3 ♂ Iran, Fars prov., Izadkhast , N 31 31 E 52 08, leg. A. Senglet ( HNHM) ; 1 ♂ Fars prov., Kazeroun, V .1953, leg. Mirzayans ( HMIM); 1 ♂ Fars prov., Kazeroun, 12. V .1967, leg. Gharib ( HMIM); 1 ♀ Fars prov., Kazeroun, Mamasani, V .1951, leg. Manuchehri ( HMIM); 1 ♂ Fars prov., Lar, 19. V .1969, leg. Pazuki & Hashemi ( HMIM); 3 ♂ 2 ♀ Fars prov., Shiraz, Kandar, Sibkhalaj, 29. VI .1963, leg. Mirzayans ( HMIM); 1 ♀ Fars prov., Kazeroun, Sarmashad, 16. V .1969; leg. Mojib ( HMIM); 4 ♂ 1♀ Fars prov., Kakan , 7.VIII.1949, leg. Mirzayans ( HMIM) ; 1 ♂ 5 ♀ Ghazvin prov. , 9.VIII.1950, leg. Mirzayans ( HMIM) ; 2 ♂ 1 ♀ Ghom prov., Lake Ghom, 27. VI .1969, leg. M.A. B ( HMIM) ; 1 ♂ 3 ♀ Ghom prov., Ghom, 900 m, 2. VI .1990, leg. Mirzayans & Abai ( HMIM); 2 ♂ 1 ♀ Hamedan prov., Asadabad , 2200m, 30.VII.1987, leg. Mirzayans & Hashemi ( HMIM) ; 3 ♂ 4 ♀ Hamedan prov., Bandarabbas, 16. V .1969, leg. Pazuki & Hashemi ( HMIM); 1 ♂ 1 ♀ Hamedan prov., Bandarabbas, Geno Mt. , 320 m, 14. VI .1992, leg. Mirzayans & Badii ( HMIM); 4 ♂ 2♀ Hamedan prov., Bandarabbas, Geno Mt. , 900 m, 8. IV .2004, leg. Moghaddam & Serri ( HMIM); 2 ♀ Hamed- an prov., Bandarabbas, Hajiabad , 1050 m, 3. V .1996, leg. Badii, Ardeh & Nazari ( HMIM); 1 ♂ Hormozgan prov., Bashagerd, Senderk , 220 m, 12. VI .1977, leg. Safavi & Pazuki ( HMIM); 1 ♂ Hormozgan prov., Minab , 23.III.1949, leg. Fasihi ( HMIM) ; 1 ♂ 4 ♀ Hormozgan prov., Minab, 12, 13. V .1969, leg. Pazuki & Hashemi ( HMIM); 2 ♂ 3 ♀ Kerman prov., Bardsir, 12. V .1952, leg. Famouri ( HMIM); 1 ♂ 1 ♀ Kerman prov., Dehbakri , 2500 m, 15. VI .1967, leg. M.A. ( HMIM) ; 1 ♂ Kerman prov., Sirjan , 6. IV .1973, leg. Abai ( HMIM); 1 ♂ 1 ♀ Kerman prov., 50 km S Sirjan , 1500 m, 2. V .1986, leg. Mirzayans & Boroumand ( HMIM); 8 ♂ 9 ♀ Kerman prov., Jebalbarez, 14. VI .1967, leg. Mirzayans & Pazuki ( HMIM); 1 ♂ 4 ♀ Kerman prov., Jiroft, 11. V .1969, leg. Pazuki & Hashemi ( HMIM); 2 ♂ 1♀ Kerman prov., Jiroft, Asminon, 11. V .1969, leg. Pazuki & Hashemi ( HMIM); 9 ♂ 9 ♀ Kerman prov., Jiroft, Mijan , 1250 m, 12. V .1968, leg. Hashemi & Pazuki ( HMIM); 3 ♀ Kerman prov., Jiroft, Mijan , 1000 m, 9. V .1969, leg. Pa- zuki & Hashemi ( HMIM) ; 1 ♂ Kermanshah prov., 12.VIII.1968, leg. Mirzayans & Mortazaviha ( HMIM) ; 1 ♀ Ker- manshah prov., Ghasre Shirin, 27. V .1961, leg. Farahbakhsh ( HMIM); 1♀ Kermanshah prov., Kerend , 1948, leg. Alexandrov ( HMIM) ; 1 ♂ Kermanshah prov., Mehran, V .1945, leg. Sarkissian ( HMIM); 2 ♂ 2 ♀ Kordestan prov., Bijar , 28.VII.1968, leg. Dezfulian & Iranshahr ( HMIM) ; 3 ♂ 1 ♀ Khuzestan prov., Aghajari, 1, 3. V .1969, leg. Aya- tollahi & Zairi ( HMIM) ; 4 ♂ 2 ♀ Khuzestan prov., Andimeshk, 8/ 9. V .1969, leg. Zairi & Ayatollahi ( HMIM); 4 ♂ 6 ♀ Khuzestan prov., Andimeshk, Bidrubeh, Ghalerezeh , 500 m, 30. IV .2001, leg. Gilasian, Hajesmailian & Mofidi ( HMIM); 1♀ Khuzestan prov., Behbahan, 11. VI .1952, leg. Vakilian ( HMIM); 7 ♂ 11 ♀ Khuzestan prov., Behbahan, 1, 2, 6. V .1969, leg. Zairi & Ayatollahi ( HMIM); 3 ♂ 1 ♀ Khuzestan prov., Behbahan, 3. VI .1969, leg. Zairi & Ayatol- lahi ( HMIM); 1 ♂ Khuzestan prov., Behbahan, Kheirabad, 3. V .1968, leg. Zairi ( HMIM); 2 ♂ 6 ♀ Kohgiluyeh & Boyerahmad prov. , Dogonbadan, 9. V .1967, leg. Zairi & Ayatollahi ( HMIM); 13 ♂ 12♀ Kohgiluyeh & Boyerahmad prov. , Dogonbadan, 8, 11. V .1968, leg. Zairi & Ayatollahi ( HMIM); 6 ♂ 1 ♀ Kohgiluyeh & Boyerahmad prov. , Do- gonbadan, 15, 28. IV .1969, leg. Ayatollahi ( HMIM); 1 ♂ Lorestan prov., Azna, Oshtorankuh, Kamandan , 2050–2300 m, 23/ 24.VIII.1994, leg. Mirzayans & Sarafrazi ( HMIM) ; 3 ♂ 1 ♀ Markazi prov., km 80 Ghom- Arak, near Easyjan, VI .1978, leg. Mash. ( HMIM); 2 ♂ 2♀ Mazandaran prov., Elborz Mts., above Rudehen , 1900–2200 m, 11. VI .2014, leg. Bologna ( MABC); 2 exx Qom prov., Neyzar, 34°16’N 50°36’E, Takht-e-Chekab ( JCRC) ; 2 ♂ 2 ♀ Persia [Raza- vi Khorasan prov.], Sabsavaran ( HNHM); 1 ex Khorasan [e- Razavi], Fazel , 9 km W Mashad ( MCNV) ; 1 ♀ Sistan & Baluchestan prov. , Bampour, 21. IV .1968, leg. Hashemi & Pazuki ( HMIM); 1 ♂ 2♀ Sistan & Baluchestan prov. , Hodar, 1. IV .1950, leg. Famouri ( HMIM); 5 ♂ 2 ♀ Sistan & Baluchestan prov. , Iranshahr, 5, 25. VI .1949, leg. Mor- salavatian ( HMIM); 1 ♂ 4 ♀ Sistan & Baluchestan prov., Iranshahr , 550 m, 25. IV .1969, leg. Hashemi & Pazuki ( HMIM); 1♀ Sistan & Baluchestan prov. , Iranshahr, 20. IV .1969, leg. Hashemi & Pazuki ( HMIM); 3 ♂ 1 ♀ Sistan & Baluchestan prov. , Iranshahr-Sarbaz, 22. IV .1968, leg. Hashemi & Pazuki ( HMIM); 5 ♂ 5 ♀ Sistan & Baluchestan prov. , Iranshahr, Damen, 24. IV .1969, leg. Hashemi & Pazuki ( HMIM); 2 ♂ 3 ♀ Sistan & Baluchestan prov. , Iran- shahr, Damen, 711. IV .1950, leg. Sarkissian ( HMIM); 4 ♂ Sistan & Baluchestan prov., Iranshahr , 25, 30.III.1951, leg. Sarkissian ( HMIM) ; 1 ♂ 4 ♀ Sistan & Baluchestan prov., Iranshahr, Hamont kouh, 1100 m, 20. IV .1968, leg. Pazuki & Hashemi ( HMIM); 1 ♂ Sistan & Baluchestan prov. , Iranshahr, 21. IV .1968, leg. Hashemi & Pazuki ( HMIM); 1 ♀ Sistan & Baluchestan prov., Iranshahr, Spake , Aliabad , 833m, N 26 47 28.9 E 60 10 46.8, 15. IV .2005 leg. Ebrahimi & Moghaddam ( HMIM); 12 ♂ 15 ♀ Sistan & Baluchestan prov., Isin , 1500 m, 17. V .1969, leg. Hash- emi & Pazuki ( HMIM) ; 7 ♂ 5♀ Sistan & Baluchestan prov., Kahnou , 800 m, 1–4. V .1969, leg. Pazuki & Hashemi ( HMIM); 1 ♂ 6 ♀ Sistan & Baluchestan prov. , Karvandar, 18. IV .1968, leg. Hashemi & Pazuki ( HMIM); 51 ♂ 47 ♀ Sistan & Baluchestan prov. , Karvandar, 22, 23. IV .1969, leg. Hashemi & Pazuki ( HMIM); 28 ♂ 23 ♀ Sistan & Bal- uchestan prov., Karvandar , 1140 m, 22–23. IV .1969, leg. Hashemi & Pazuki ( HMIM); 8 ♂ 9 ♀ Sistan & Baluchestan prov. , Karvandar, Alidar, 24. IV .1969, leg. Hashemi & Pazuki ( HMIM); 3 ♀ Sistan & Baluchestan prov., Karvandar, Alidar , 800–1000 m, 1. V .1969, leg. Pazuki & Hashemi ( HMIM); 1 ♂ Sistan & Baluchestan prov., Karvandar , Iran- shahr, 16.VII.1972, leg. Abai ( HMIM) ; 1 ♀ Sistan & Baluchestan prov. , Pishin, 17. IV .1950, leg. Famouri ( HMIM); 4 ♂ 4 ♀ Sistan & Baluchestan prov. , Pishin, 22. V .1950, leg. Famouri ( HMIM); 1♀ Sistan & Baluchestan prov., Negor , 7.III.1997, leg. Mirzayans, Badii & Sarafrazi ( HMIM) ; 1 ♂ 2 ♀ Sistan & Baluchestan prov. , Saravan, Bam- posht, 26. IV .1950, leg. Mirsalavatian ( HMIM); 3 ♂ 6 ♀ Sistan & Baluchestan prov., Sarbaz , 1200 m, 23. IV .1968, leg. Pazuki & Hashemi ( HMIM); 10 ♂ 7 ♀ Sistan & Baluchestan prov. , Sarbaz- Rask, 24. IV .1968, leg. Hashemi & Pazuki ( HMIM); 2 ♂ 3 ♀ Sistan & Baluchestan prov. , Sarbaz-Zangian, 13. IV .1949, leg. Mirsalavatian ( HMIM); 2 ♂ 2 ♀ Sistan & Baluchestan prov. , Rask, 24. IV .1968, leg. Pazuki & Hashemi ( HMIM); 1 ♂ Sistan & Baluchestan prov., Tang-e Sarhe Nikshahr , 23. III. 1949, leg. Sharif ( HMIM) ; 3 ♀ Sistan & Baluchestan prov. , Tangesarhe, 10. IV .1973, leg. Safavi & Broumand ( HMIM); 1 ♀ Sistan & Baluchestan prov. , Nikshahr, 23. IV .1968, leg. Hashemi & Pazuki ( HMIM); 2 ♂ Sistan & Baluchestan prov. , Nikshahr, 28. IV .1969, leg. Pazuki & Hashemi ( HMIM); 16 ♂ 13 ♀ Sistan & Baluchestan prov., Tang-e Sarhe , 1000 m, 27, 30. IV .1969, leg. Pazuki & Hashemi ( HMIM); 1♀ Sis- tan & Baluchestan prov. , Zabol, 5. VI .1949, leg. Mirzayans ( HMIM); 2 ♂ Sistan & Baluchestan prov. , Zabol, 25. V .1950, leg. Mirzayans ( HMIM); 1 ♂ Sistan & Baluchestan prov. , Zahedan, Nosratabad, 21. IV .1969, leg. Pazuki & Hashemi ( HMIM); 1 ♂ 1 ♀ Persien, Baluchistan, IV . 1949, leg. Sharif ( HNHM); 1 ♂ Beluchistan, nrdwestl. Iranshahr ( Trocken Tal ), 15.III.1954, leg. Richter & Schäuffele ( HNHM) ; 2 ♂ [Semnan prov.], Shrud, 9. V .1956, leg. Schmid ( HNHM); 1 ex Semnan prov., 5 km NE Agareh ( TGC) ; 1 ♂ 2 ♀ Tehran prov., Abshar , 10.VII.1947, leg. Mirsalavatian ( HMIM) ; 1 ♂ 1 ♀ Tehran prov., Evin , 18.VII.1948 leg. Mirzayans ( HMIM) ; 8 ♂ 14 ♀ ( HMIM) ; 1 ♀ ( HNHM) Tehran prov., Evin , 6. VI .1965, leg. Mirzayans & Daneshpazhouh; 4 ♂ 2 ♀ ( MABC) , 2 ♂ 2♀ ( HNHM) Tehran prov., Golhak , bei Teheran, 1700 m, 9–23. VI .1961, leg. Klapperich; 1 ♂ Tehran prov., Lashgarak , 1650 m, 18. VI .2004, leg. Serri & Frisch ( HMIM); 1 ♂ 1 ♀ Persien, Tehran, Mahmudieh , steppe, 15.VII.1948, coll. P. Allen ( HNHM) ; 1 ♂ Tehran prov., Marounak , 12.VIII.1948, leg. Vaezi ( HMIM) ; 1 ♀ Tehran prov., Pachalak, 29. VI .1948, leg. Mirzayans ( HMIM); 1 ♂ Tehran prov., Sohanak, 16. VI .1998, leg. Zareiee & Razi ( HMIM); 1 ♂ 1 ♀ Tehran prov., Tehran, 11.VIII.1947, leg. Mirsalavatian ( HMIM); 65 exx. Tehran, Golhak ( MSNV) ; 1 ♂ Tehran prov., Vara- min, VI .1947, leg. Alexandrov ( HMIM); 1 ♂ 1 ♀ Tehran prov., Varamin, IV .1950, leg. Eghlidi ( HMIM); 1 ♀ Yazd prov., Abarghou, Aliabad, 25. VI .1972, leg. Safavi ( HMIM); 1 ♂ Strauss Persien ( HNHM) ; 1 ♂ Persia , 1862–63, coll. Soloris, R . Oberthür, Eing. Nr. 4, 1956 ( HNHM) . Iraq. 3 ♂ 6 ♀ Assur, Mesopotamia , coll. Hauser ( HNHM) ; 1 ♀ Assur, Mesopotamia, V .1908 ( HNHM); 1 ♂ Assur, Mesopotamia, Exp. Nat. O. V ., 1910, leg. Pietschmann ( HNHM); 1 ♂ Mesopotamia, Mosul, V .1909, coll. Hauser ( HNHM); 1 ♂ Tekrit, 7. V .1910, Mesopot. Exp. Nat. O. V. ( HNHM) . Pakistan. 1 ex., NWFP , 85 km NE Chitral, Chitral-Mastuj road ( VVC) . Turkmenistan. 1 ♂ coll. Reitter ( HNHM) ; 2 ♀ Kopet Dag Mt. , 10 km S Aldere, N38 14 E 56 46, 600 –1000 m, 27. VI .1992, leg. G. Fabian ( HNHM); 1 ♂ Kopet Dagh, Germab , 1894, coll. Hauser ( HNHM) ; 3 ♂ 1 ♀ Chuli, 20. VI .1963, leg. Kozolvo ( MABC).

Records from literature. The taxonomy of this species is also very confused, so we prefer to consider only confirmed or congruent records from the literature. Afghanistan. Afghanistan ( Kaszab 1968 as javeti ); Darreh-Gazak ( Kaszab 1973); Hérat ( Kaszab 1961 as javeti and ab. heratensis nov.); Nuristan, Bashgultal, Parigil-Patscheigil ( Kaszab 1958, 1973 both as javeti umbilicata); near Kabul ( Kaszab 1958, typ. loc., 1973 both as javeti umbilicata); Kabul West ( Kaszab 1965); Nuristan (Borchman 1936); E Afghanstan, Neugrahar, Darunta ( Kaszab 1973 as javeti umbilicata); Borgh, Logar ( Kaszab 1973 as javeti umbilicata); Qal’eh Lakou nr. Kabul valley ( Kaszab 1973 as javeti umbilicata); Kouh-Djaouz, nr. Tang-Saldan 20 km W Kaboul ( Kaszab 1973 as javeti umbilicata); Khairou Khil, 30 km NW Djelalabad ( Kaszab 1973 as javeti umbilicata); Laghman, Samakat ( Kaszab 1981 as javeti umbilicata); Ghazni, 50 km NW Moqur ( Kaszab 1981 as javeti umbilicata); Nangarhar, Dast-e-Gamberay ( Kaszab 1981 as javeti umbilicata); Kabul-Sarobi ( Kaszab 1981); Parigil and Patscheigil ( Borchmann 1936 as colligata ). Iran. Südpersien ( Kollar & Redtenbacher 1850 type locality); Iran (also as Perse) ( MSNV; Marseul 1870, 1872; Sumakov, 1915, 1930; Mader 1927a; Kuzin 1954; Kaszab 1968; Bologna 2008; Fekrat & Modarres Awal 2012; Bologna et al 2018; Ghahari & Campos-Soldini 2019 all as javeti ); S Iran (Redtenbacher 1850 type locality); Fars ( Koçak & Kemal 2015); Bushehr, Fars, Khuzestan, Kohgiluyeh & Boyerahmad ( Serri 2015); Fars, Hormozgan, Kerman, Kermanshah, Kordestan, Khuzestan, Ghazvin, Sistan & Baluchestan, Zanjan ( Mirzayans 1970; Modarres Awal 1997, 2012; Ghahari & Campos-Soldini 2019, all as javeti ); Esfahan, W Ná-in ( Mohammad-Momeni et al. 2016 as javeti ); Esfahan ( Kaszab 1968; Ghahari & Campos-Soldini 2019 as javeti ); Hamedan, Nahavand Co. ( Nikbakhtzadeh & Tirgari 2002 as Mylabris schreibersi ); 100 km N Iranshar ( Kaszab 1968 as javeti ); Khorasan, Razavi prov., Mashhad ( Fekrat & Modarres Awal 2012 also as javeti ); S Khorasan Birjand ( Fekrat & Modarres Awal 2012 also as javeti ); Kopet dag ( Kuzin 1954 as javeti ); Markazi ( Faraji et al. 2012; Moslemi et al. 2015 both as javeti ); Shiraz ( Sumakov 1930); Sistan & Baluchestan ( Kaszab 1968 as javeti ) Tehran ( Mirzayans 1970; Modarres Awal 1997, 2012; Ezzoddinloo & Mohammadian 2012 as javeti ). Iraq. Iraq ( Derwesh 1965 as javeti ; Fekrat & Modarres Awal 2012 as javeti ; Bologna et al. 2018 as javeti ; Ghahari & Campos-Soldini 2019 as javeti ); Assur ( Holdhaus 1920).

Transcaucasia. Transcaucasia ( Faldermann 1837, 1839 as superba ). «Turkestan». ( Dokhtouroff 1889; Sumakov, 1915; Bologna 2008; Fekrat & Modarres Awal 2012; Ghahari & Campos-Soldini 2019 all as javeti ). Turkmenistan. Turkmenia ( Sumakov 1915; Mader 1927a; Bologna 2008; Fekrat & Modarres Awal 2012; Bologna et al 2018; Ghahari & Campos-Soldini 2019 all as javeti ); Transcaspia ( Mader 1927a; Kaszab 1968 both as javeti ); Askhabad ( Reitter 1889; Sumakov 1930 as javeti ); Kopet Dag ( Kuzin 1954 as javeti ); Askhabad and Tekke ( Dokhtouroff 1889 as javeti ); Samaná Syr ( Heyden & Kraatz 1883 as javeti ); Nokhur; Arman saad-Kizil arvat; Gaudan; Kaaşka; Firyuza; Tejen ( Reichardt 1934 as javeti ).

Description. Body length 7–24 mm. Body black, elytra yellowish or reddish brown, sometimes dark red, with black pattern. Head subquadrate; antennomere III less than 1/4 time longer than IV, V–VII transverse and widest, antennomeres VII–IX progressively narrowed, male XI ( Fig. 55 ) elongated apically, spindle shaped; in female last antennomere short ( Fig. 56 ). Pronotum ( Fig. 17 ) as long as wide or slightly longer, with anterior and middle depression, one middle longitudinal furrow, distinct punctures; mesosternun as in Fig. 61 ; each elytron with two black spots on anterior third, one wide middle fascia, and one black fascia on apical part ( Fig. 62 ). Last male ventrite convex, except one narrow depression in middle, posterior margin deeply emarginated in middle; aedeagus ( Figs 59–60 ) with endophallic hook slightly narrow, distal aedeagal hook small and curved, very close to proximal one and both almost at apex, proximal hook short and nearly straight; gonoforceps ( Figs 57–58 ) narrow and straight in noth ventral and lateral view.

Taxonomic remarks. As explained in the taxonomic remarks of H. cingulatus , after checking the syntypes of both Mylabris colligata and Mylabris javeti (MNHN, NHMW) we concluded that these species are synonyms and colligata has the priority of name.

We propose the following taxonomic act: Mylabris javeti Marseul, 1879 = Hycleus colligatus (Redtenbacher, 1850) new synonymy.

Kaszab (1958) described Mylabris javeti umbilicatus as a subspecies endemic to Afghanistan, and the variety subcordata . This subspecies differs from the nominate form mostly by its dense and robust punctation. Checking some hundred specimens of this species from the whole range, we found this variation also among some populations of Iran, sympatric with the nominate form, so we consider this subspecies only as a variation without taxonomic value. Apart from the pronotal punctures, the shape of the last antennomere of males and aedeagal hooks show intrapopulation variations ( Fig. 62 ). The molecular study of this complex confirms this huge variation.

We propose the following taxonomic act: Mylabris javeti umbilicatus Kaszab, 1958 = Hycleus colligatus Redentbacher, 1850 new synonymy.

Distribution. This species is widespread in Iran and in northern Afghanistan, and marginally distributed in the Transcaucasian area, Iraq, Northwest Pakistan and Turkmenistan.

Figures

FIGURES 55–62. Hycleus colligatus (Redtenbacher): 55. male antenna; 56. female antenna; 57. tegmen, ventral view; 58. tegmen, lateral view; 59. aedeagus, lateral view; 60. variations of aedeagus; 61. mesosternum; 62. elytral pattern. Scale bar: 1 mm.

FIGURES 13–24. Head and pronotum of: 13. Hycleus apicipennis; 14. H. balteatus; 15 H. bipunctatus; 16. H. cingulatus; 17. H. colligatus; 18. H. hayki; 19. H. lindbergi; 20. H. pirata; 21. H. schauffelei; 22. H. sexmaculatus; 23. H. tenuepictus; 24. H. trianguliferus.