Hoploparia biserialis Fritsch

Tshudy, Dale M., Hyžný, Matúš, Veselská, Martina Kočová & Jagt, John W. M., 2022, Taxonomic revision of the extinct clawed lobster genus Oncopareia Bosquet, 1854 (Decapoda, Astacidea, Nephropidae), Palaeontologia Electronica (a 20) 25 (2), pp. 1-31 : 21-22

Hoploparia biserialis Fritsch in Fritsch and Kafka, 1887

Figure 11A–C

Partim 1887 Hoploparia biserialis Fritsch in Fritsch and Kafka, p. 35, text-fig. 56, pl. 3, fig. 5; pl. 5, fig. 1 [non pl. 5, fig. 2 = Oncopareia bredai Bosquet, 1854 ]

1929 Hoploparia biserialis Fritsch in Fritsch and Kafka; Glaessner, p. 216.

1941 Hoploparia “biserialis ” Fritsch in Fritsch and Kafka; Mertin, p. 176, text-fig. 7h.

1941? Oncopareia biserialis (Fritsch in Fritsch and Kafka); Mertin, text-fig. 8a.

2010 Hoploparia biserialis Fritsch in Fritsch and Kafka; Schweitzer et al., p. 28.

2010 Oncopareia biseralis [sic] Fritsch in Fritsch and Kafka; Schweitzer et al., p. 31.

2018 Hoploparia biserialis Fritsch in Fritsch and Kafka; Tshudy et al., p. 182.

Material examined. Four specimens in the original

Fritsch Collection from the National Museum in

Prague, labelled as Hoploparia biserialis , including

NM O4044 (Orig. #91; cephalothorax and pleon),

NM O3470 (Orig. #86; cephalothorax), NM O3457

(Orig. #87; isolated cutter claw) and NM 06861 (cephalothorax) . NM O4044 ( Figure 11A ) is herein designated lectotype of the species .

Diagnosis. Hoploparia with combined postcervical and branchiocardiac grooves extending to hepatic groove. Antennal carina missing. Cutter claw elongate, with fingers longer than palm.

Remarks. Two specimens, NM O3470 and NM O6861, representing isolated cephalothoraxes of O. bredai were removed from the type collection of H. biserialis (see above). The remaining specimens, NM O4044 and NM O3457, confidently confirm placement in the genus Hoploparia . The long, pointed pleura of H. biserialis NM O 4044 (see Fritsch and Kafka, 1887, text-fig. 56; Figure 11A here) are clearly Hoploparia - or Homarus -like and very unlike those of Oncopareia . No claw pairs are known from this specimen. The isolated cutter claw NM O3457 (e.g., Fritsch and Kafka, 1887, pl. 3, fig. 5; Figure 11B–C here) is Hoploparia -like; with a subtriangular palm, the inner margin of which has numerous large, anteriorly directed spines in two rows; fingers are long and robust, lack dentition on occlusal margins and become narrower distally.

It is important to note, that Mertin (1941) considered these two specimens of H. biserialis (sensu Fritsch and Kafka, 1887) , NM O3457 and NM O4044, to belong to different taxa. The isolated cutter claw NM O3457 ( Mertin, 1941, text-fig. 7h), in Mertin’s (1941) view closely resembling that of H. longimana , was referred to as Hoploparia “biserialis ”, whereas the carapace with pleon NM O4044 ( Mertin, 1941, text-fig. 8a) was treated as? Oncopareia biserialis . This pleon with cordate, pointed pleura, and cephalothorax with smooth rostrum did not deter Mertin (1941, p. 181) from placing the species in Oncopareia , which, by his definition, possesses a “stark bedornten” (strongly spiny) rostrum. Although the material comes neither from the same locality nor the same lithostratigraphic unit, we here refer both these specimens to Hoploparia biserialis , following Fritsch and Kafka (1887).

Range and occurrence. Upper Cretaceous, lower–middle Turonian of the Czech Republic; NM O3457 comes from the middle Turonian (lower part of the Jizera Formation) at Bechlín near Roudnice nad Labem and NM O4044 is from the lower–middle Turonian (Bílá Hora Formation) of Bílá Hora (‘Weissenberg’) in Prague.

Figures

FIGURE 11. Hoploparia biserialis Fritsch, in Fritsch and Kafka, 1887; lower–middle Turonian (Bílá Hora Formation) in Prague, Czech Republic. A, Cephalothorax with pleon and tailfan in lateral view (lectotype, NM O4044). B-C, Isolated right cutter claw in dorsal (B) and lateral (C) views (NM O3457). Specimen in A was whitened with ammonium chloride prior to photography.