Heterostegina reticulata multifida ( Bieda, 1949 )

Less, György, Özcan, Ercan, Papazzoni, Cesare A. & Stockar, Rudolf, 2008, The middle to late Eocene evolution of nummulitid foraminifer Heterostegina in the Western Tethys, Acta Palaeontologica Polonica 53 (2), pp. 317-350 : 335-336

Heterostegina reticulata multifida ( Bieda, 1949)

Fig. 12J–M, O–Q.

part 1949 Grzybowskia multifida sp. nov.; Bieda 1949: 153–158, 168–

173, pl. 3: 1, 3, 7; pl. 4: 1. (non 2). part 1978 Heterostegina reticulata multifida ( Bieda, 1949) ; Herb 1978:

758, text−figs. 14–17 (non 11). part 1993 Heterostegina reticulata Rütimeyer, 1850 ; Papazzoni and

Sirotti 1993: 532, 535–536, pl. 1: 1; pl. 2: 1, 2. 1994 Heterostegina reticulata Rütimeyer, 1850 ; Papazzoni 1994: pl.

2: 1. 1999 Heterostegina reticulata Rütimeyer, 1850 ; Romero et al. 1999:

87, text−figs. 10–11.

Diagnosis.— Heterostegina reticulata populations with X mean ranging from 7.2 to 11.

The equatorial section of A−forms.—The shape of the chamberlets is close to hexagonal; their arrangement is almost regular. For numerical features see Table 5.

Remarks.—According to Bieda (1949), the type locality of Grzybowskia multifida is a pebbly sandstone (“grès conglomeratique”) intercalation in the Podhale flysch in Szaflary, in the northern slope of the Polish Tatras. It is associated with characteristic middle Eocene forms like large−sized Nummulites of the “ perforatus ”, “ millecaput ”, and “ brongniarti ” groups and Assilina exponens on the one hand and also with upper Eocene forms such as Nummulites fabianii and several small radiate Nummulites . Both the sedimentary features and the composition of the fossil assemblage indicate the mixing of the latter caused by redeposition. Bieda (1949) has provided three good photos of the new species from the type locality. They seem to confirm the mixing of the fauna since the form in his pl. 4: 2 is much more advanced than the other two as recognized already by Herb (1978). Bieda (1949) marked out the specimen in pl. 3: 1 as typical for the equatorial section of A−forms. In this specimen the parameter X counted by us is 8, therefore we apply the name “ multifida ” to populations with the above diagnosis. X mean of the four specimens from the upper Hohgant sandstone of Elsigen ( Switzerland) illustrated by Herb (1978) is 9.75, which is consistent with the above diagnosis although with some reservations because of the few numbers of specimens. The forms illustrated by Papazzoni and Sirotti (1993) and by Papazzoni (1994) are from the vicinity of our sample Mossano 2, therefore they are tentatively identified with “ multifida ”. The specimens figured by Romero et al. (1999) from Puig Aguilera are re−evaluated in this paper and according to the parameters classified in synonymy with the present subspecies.

Geographic and stratigraphic distribution.—Latest Bartonian (SBZ 18 C) of Italy (samples Mossano 2 and Verona, Monte Cavro 4), Switzerland (Elsigen), NE Spain (Puig Aguilera) and Poland (Szaflary, Podhale Depression).

Heterostegina reticulata helvetica Kaufmann, 1867 Figs. 12N, R, S, 13A–L.

1867 Heterostegina helvetica sp. nov.; Kaufmann 1867: 153, pl. 9: 6–10.

part 1978 Heterostegina reticulata reticulata Rütimeyer, 1850 ; Herb 1978: 758–759, text−figs. 7–10, 12–13, 35.

1999 Heterostegina reticulata reticulata Rütimeyer, 1850 ; Stockar 1999: 6, 9, pl. 1: 4–5; pl. 3: 1–6.

2004 Heterostegina reticulata urhidensis ssp. nov.; Less and Gyalog 2004: pl. (black and white) 1: 5.

Diagnosis.— Heterostegina reticulata populations with X mean ranging from 4.4 to 7.2.

The equatorial section of A−forms.—The shape of the quite regularly arranged chamberlets is hexagonal. For numerical features see Table 5.

Remarks.—The biometrical parameters of Heterostegina reticulata from sample Gschwänt, the type locality of Kaufmann’s (1867) H. helvetica , designated by Herb (1978), indicate that populations with the above definition belong to this subspecies and clearly differ from those of H. reticulata reticulata with type locality in Oberbergli. Therefore, the name “ helvetica ” is valid in the subspecific level despite that Herb (1978) put Kaufmann’s (1867) H. helvetica in synonymy with Rütimeyer’s (1850) H. reticulata on both the specific and subspecific levels. Spiroclypeus could not be found in Gschwänt, therefore the early Priabonian age of this locality, attributed by Herb (1978), cannot be confirmed. The forms illustrated by Stockar (1999) are from our sample Prella 4. This population is re−evaluated and based on its parameters, it is ranked in synonymy with the present subspecies. Less and Gyalog (2004) introduced the name “ urhidensis ” for Heterostegina reticulata populations from samples Úrhida 4 to 7 with no description. After the revision of H. r. helvetica from its type locality the new name turned out to be invalid.

Geographic and stratigraphic distribution.—Latest Bartonian (SBZ 18 C) of Hungary (Úrhida, samples 4 to 8), Switzerland (Gschwänt; sample Prella 4), and Armenia (Biralu, “ millecaput ” beds).

Heterostegina reticulata reticulata Rütimeyer, 1850 Figs. 13M–R, 14A–F .

1850 Heterostegina reticulata sp. nov.; Rütimeyer 1850: 109, pl. 4: 61.

part 1978 Heterostegina reticulata reticulata Rütimeyer, 1850 ; Herb 1978: 758–759, text−figs. 2–6 (non 7–10, 12–13, 35).

2004 Heterostegina reticulata reticulata Rütimeyer, 1850 ; Less and Gyalog 2004: pl. (black and white) 2: 2.

Diagnosis.— Heterostegina reticulata populations with X mean ranging from 2.8 to 4.4.

The equatorial section of A−forms.—The shape of the regularly arranged chamberlets is distinctly to slightly hexagonal. For numerical features see Table 5.

Remarks.—The biometrical parameters of Heterostegina reticulata from sample Oberbergli, the type locality of Rütimeyer’s (1850) H. reticulata , designated by Herb (1978), indicate that populations with the above definition belong to this subspecies. Spiroclypeus could not be found in Oberbergli, therefore the early Priabonian age of this locality, attributed by Herb (1978), cannot be confirmed. About the distinction of “ helvetica ” from “ reticulata ” put in synonymy by Herb (1978) see remarks on the former.

Geographic and stratigraphic distribution.—Latest Bartonian (SBZ 18 C) of Italy (Mossano, sample 3), Hungary (Úrhida, sample 9), Switzerland (Oberbergli) and Armenia (Vedi, “ millecaput ” beds).

Figures

Fig. 14. Latest Bartonian and early Priabonian Heterostegina from different Western Tethyan localities. A–F. Heterostegina reticulata reticulata Rütimeyer, 1850, latest Bartonian, SBZ 18 C. A–E. Mossano 3 (N Italy), equatorial sections, MÁFI E. 9549, A−form (A), MÁFI E. 9550, A−form (B), MÁFI E. 9551, A−form (C), MÁFI E. 9552, B−form (D), MÁFI E. 9553, A−form (E). F. Vedi (Armenia), MÁFI E. 9554, A−form, equatorial section. G–R. Heterostegina reticulata mossanensis ssp. nov., earliest Priabonian, SBZ 19 A. G–J. Mossano 6, equatorial sections, holotype, MÁFI E. 9555, A−form (G), paratype, MÁFI E. 9556, A−form (H), paratype, MÁFI E. 9557, A−form (I), paratype, MÁFI E. 9558 (J), B−form. K–M. Úrhida 10 (Hungary), Ą