Hadrodontina aequabilis Staesche, 1964

Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas, 2022, A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China, Swiss Journal of Palaeontology (19) 141 (1), pp. 1-61 : 53-56

publication ID

https://doi.org/ 10.1186/s13358-022-00259-x

DOI

https://doi.org/10.5281/zenodo.13128558

persistent identifier

https://treatment.plazi.org/id/038B1D1D-645D-FF86-FCA1-E85BD2E2FBC7

treatment provided by

Felipe

scientific name

Hadrodontina aequabilis Staesche, 1964
status

 

Hadrodontina aequabilis Staesche, 1964

P 1 elements: Figs. 27F–M View Fig ; 28A, C, E View Fig

P 2 View Fig elements: Figs. 27B–E View Fig ; 28B, D View Fig

S 2 View Fig elements: Fig. 27A View Fig

P 1 View Fig element:

*1964 Hadrodontina aequabilis n. sp.; Staesche, p. 275, figs. 43, 44.

1984 Sweetocristatus unicus n. sp.; Dagis, pp. 37–38, pl. X, figs. 6–9.

1990 Pachycladina peculiaris n. sp.; Shunxin (Zhang), pl. 2, fig. 4

1991 Pachycladina peculiaris n. sp.; Zhang in Zhang & Yang, p. 40, pl. 3, figs. 1, 2.

1997 Parapachycladina peculiaris Zhang ; Shunxin et al., pp. 65–69, pl. 1, figs. 1–2, pl. 2, figs. 1–2, pl. 3 figs. 1–2.

2004 Ellisonia sp. aff. E. triassica Müller, 1956 ; Koike et al., figs. 8.7, 8.8.

2009 Ellisonia? cf. peculiaris Sweet ; Igo in Shigeta et al., p. 182, fig. 152.22.

2013 Parachirognathus geiseri Clark ; Yan et al., p. 516, fig. 6 FF.

2015 Parachirognathus peculiaris Zhang & Guo ; Chen et al., figs. 8.20–21, 24.

2015 Sweetocristatus unicus Dagis ; Chen et al., fig. 8.23. 2016 Hadrodontina aequabilis Staesche ; Koike, pp. 164–167, fig. 2, nr. 1–3.

2018 Hadrodontina aequabilis Staesche ; Maekawa in Maekawa et al., p. 18, figs. 13.3?–13.4 (only).

2020 Hadrodontina aequabilis Staesche ; Sun et al., figs. 1–7 (natural assemblage of the Hadrodontina apparatus).

P 2 element:

*1964 Hadrodontina aequabilis n. sp.; Staesche, p. 275, fig. 44.

2004 Ellisonia sp. aff. E. triassica Müller ; Koike et al., p. 247, fig. 8.6

2016 Hadrodontina aequabilis Staesche ; Koike, p. 165, figs. 2.4 (P2 element).

2020 Hadrodontina aequabilis Staesche ; Sun et al., figs. 1–7 (natural assemblage of the Hadrodontina apparatus).

S 2 element. 2004 Ellisonia sp. aff. E. triassica Müller ; Koike et al., p. 247, fig. 8.4

2016 Hadrodontina aequabilis Staesche ; Koike, p. 165, figs. 2.9–11.

2020 Hadrodontina aequabilis Staesche ; Sun et al., figs. 1–7 (natural assemblage of the Hadrodontina apparatus).

Material. P 1, more than 40; P 2, more than 40; S 2 more than 10.

Revised diagnosis. Robust to slender angulate P 1 element. Te 6–8 (usually 7) denticles are rather short, radiating, conical, with subtriangular free ends and they are increasingly reclined towards the posterior. Te concave lower side is occupied by a large and wide groove.

Remarks. Te observed variation in the denticulation of the P 1 and P 2 elements suggests there may be scope for further specific differentiation: in some elements the height of the denticles changes smoothly along the unit ( Fig. 28B View Fig ), whereas in others the denticles are alternatively high and low ( Fig. 28D View Fig ). Dagis (1984) assigned some specimens (here considered as junior synonyms of Ha. aequabilis ) to Sweetocristatus unicus . Yet, the genus Sweetocristatus Szaniawski (in Szaniawski & Malkowski, 1979) was established for Upper Artinskian to Lower Guadalupian (Permian) P 1 elements that, although superficially similar to homologous elements of Hadrodontina and Pachycladina , bear a more developed, higher cusp, a higher carina and have a more elongated process. Te cusp in Ellisonia P 1 elements is more conspicuous than in those of Hadrodontina or Pachycladina . Chen et al. (2015) illustrated coeval specimens from Jiarong they assigned to either Sweetocristatus unicus or Parachirognathus peculiaris (see their figs. 8.20 and 8.23) but we fail to distinguish them from Hadrodontina aequabilis . With three natural assemblages of Hadrodontina aequabilis from the late Smithian Helongshan Formation, South China, Sun et al. (2020) observations fits with our findings; the angulate shape and no distinct cusp in the P 1 element and the distally twisting shape of the P 2 element.

Occurrence. Russia; Smithian Hedenstroemia and tardus zone, Siberia ( Dagis 1984), Zhitkov Formation, Smithian age, Abrek Bay area, South Primorye ( Shigeta, 2009). China; Beisi Formation, Taiping, Pingguo Western Guangxi Province ( Shunxin et al., 1997), Luolou Formation, Smithian age, Jiarong and Bianyang, Nanpanjiang Basin, southern Guizhou Province ( Chen et al., 2015; Yan et al., 2013), Late Smithian Helongshan Formation, Chaohu, Anhui Province ( Sun et al., 2020). Japan: Taho Formation, Shirokawa-cho, Higashiuwa-gun, Ehime Prefecture (below Smithian–Spathian boundary) ( Koike 2016; Koike et al., 2004). Europe: Campiller member, Werfen Formation, Skyth, South Tirol, Italy ( Staesche, 1964), Slovenia ( Kolar-Jurkovšek & Jurkovšek, 2019).

Kingdom

Animalia

Phylum

Chordata

Order

Ozarkodinida

Family

Ellisoniidae

Genus

Hadrodontina

Loc

Hadrodontina aequabilis Staesche, 1964

Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas 2022
2022
Loc

Hadrodontina aequabilis

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Sweetocristatus unicus

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Pachycladina peculiaris

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Pachycladina peculiaris

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Hadrodontina aequabilis

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Hadrodontina aequabilis

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Hadrodontina aequabilis

Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022
2022
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina

Staesche 1964
1964
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina

Staesche 1964
1964
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina aequabilis

Staesche 1964
1964
Loc

Hadrodontina

Staesche 1964
1964
Loc

Ellisonia sp. aff. E. triassica Müller, 1956

Muller 1956
1956
Loc

Ellisonia? cf. peculiaris Sweet

Muller 1956
1956
Loc

E. triassica Müller

Muller 1956
1956
Loc

E. triassica Müller

Muller 1956
1956
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