Galium, K. Schum.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1976, Flora Europaea. Volume 4. Plantaginaceae to Compositae (and Rubiaceae), Cambridge University Press : 20

Sect. Trachygalium K. Schum. Perennial herbs with slender rooting rhizomes and subterranean stolons. Stem 4-angled, retrorsely aculeolate and rough on the angles. Leaves in whorls of 6-10, shortly awned, thinly coriaceous, shiny above, 1 -veined, the margins antrorsely and retrorsely scabrid. Inflorescence ovoid or cylindrical, lax, the ultimate branches usually ebracteate. Corolla infundibuliform or cup-shaped, rarely rotate, white, reddish or greenish-yellow; lobes acute to shortly apiculate, usually papillose. Fruit mostly granulate and glabrous or hairy.