Fannia losgateados, Dominguez, 2007

Dominguez, Martha Cecilia, 2007, A taxonomic revision of the Southern South American species of the genus FANNIA Robineau-Desvoidy (Diptera: Fanniidae), Papéis Avulsos de Zoologia (São Paulo) 47 (24), pp. 289-347 : 323-325

Fannia losgateados sp. n.

Figs. 13 A-F

Diagnosis: Small dark species. Frontal vitta at narrowest point 1.9x width of anterior ocellus. Eye bare. Parafacial bare. Antenna and palpus black. Scutum dark brown to black, with broad dark brown shining vitta along space between dc and ia lines. Two very short and thin pra near spal. T 2 constricted at base, with a weak subbasal protuberance and 3 submedial a. C3 with three p at apex of posterior margin. F3 thin, with a preapical protuberance 1.5x width of femur, on posteroventral surface, ventral and posteroventral surfaces covered with long hair-like setae, 1.5x femoral width and a tuft of long pv, 2x femoral width, coinciding with the posteroventral protuberance. T 3 with 3 to 4 medial ad. Wing clear with brown tint. Halter yellow. Abdomen heart-shaped, dark brown to black, base and lateral margins of tergites 3, 4, and 5 light blue-greyish.

Etimology: The species name is an apposition that refers to Los Gateados ravine, in Potrerrillos , province of Mendoza, where the holotype was collected .

Type material examined: Holotype o: Argentina: Mendoza, Potrerillos, Quebrada de los Gateados , X-2001, col. Domínguez ( IADIZA) . Paratypes: 4 n, same data as holotype ( IADIZA); 1 o, Quebrada Aguas Buenas, XII-2000, cols. Domínguez & Roig ( IADIZA) .

Description

Male: body length 3.5-3.8 mm.

Head: Frontal vitta black, at narrowest point 1.9x width of anterior ocellus. Fronto-orbital plate silvery pruinose, at uppermost fr lightly narrower than width of anterior ocellus. Twelve long fr, 1.4x length of flagellum. Eye bare; pocl uniform in length. Facialia dark grey. Parafacial silvery, at base of flagellum 0.7x width of same and bare. Facial carina black. Gena dark grey. Scape and pedicel black, flagellum 2 to 2.25x as long as wide, heavily dark grey pruinose. Arista completely black, short pubescent (pubescence shorter than aristal width at base). Palpus black and spatulated, the apex more than 2x width of base. Lower oral margin slightly concave and grooveless.

Thorax: Scutum dark brown to black, dark brown shining vitta along space between dc and ia lines; pleura and apex of scutellum dark grey; 3+3 acr and 1 precutellar acrs; 2+3 dc; pprn covered with setulae; 2 very short and thin pra near spal, almost indistinguishable from ground setulae; prealar area covered with setulae; 2 prepm surrounded by numerous setulae.

Legs: Dark brown, tarsi black. F1 normal to thin (length/width = 5.5), with 1 row of pd and 2 rows of p as long as femoral width, the most ventral row of p is hair-like; 1 row of pv short and spaced at base, as long as femoral width at apex. T 1 with a subapical long pd; one apical v and pv. Fore tarsomere 1 0.3x length of t1, tarsomere 4 as wide as long. F2 with 1 row of ad and 1 row of a; 1 row of av, spaced at base and grouped at apex; 1 long basal pv and a complete row of pv that is doubled at apex with a second hooked row; 1 row of p, longer and ventrally directed towards apex. T 2 constricted at base, with a weak subbasal protuberance; ventral pubescence short, shorter than 0.5x width of T 2; 3 submedial and 3 apical a; 1 subapical ad; 1 apical av; 2 submedial and 2 apical pd; 1 apical pv; apical p absent. C3 with 3 p at apex of posterior margin. F3 thin (length/width = 8), with a preapical posteroventral protuberance, 1.5x femoral width; 1 row of ad longer and dorsally displaced towards apex; 3 row of short a, 2 occupying basal half, third complete; 2 rows of av, the most dorsal as long as femoral width, the ventral row short and thin; ventral and posteroventral surfaces covered with long hair-like setae, 1.5x femoral width; tuft of long pv, 2x femoral width, coinciding with protuberance ( Fig. 13A ). T 3 with 1 submedial and 1 subapical d; 3 to 4 medial ad, apical ad absent; 1 apical and 5 subapical av; apex of ventral and posteroventral surfaces with a strong ctenidium ( Fig. 13A ).

Wing: Clear with brown tint. Lower calypter slightly oval, upper rounded, both yellowish-white and dark brown in outer margin. Knob of halter yellow, base light brown.

Abdomen: Heart-shaped, tergite 2 2x width of tergite 4; dark brown to black, base and lateral margins of

tergites 3, 4 and 5 light blue-greyish. Hypopygium not projecting.

Postabdomen: Posterior margin of sternite 5 bilobated, with scarce setae on central line, numerous on posterior margin ( Fig. 13B ). Lateral margins of hypandrium grooved; hypandrial arms expanded at apex and directed outwards; postgonite inconspicuous; aedeagus membranous ( Fig. 13C ). Epandrium anteriorly broadened, bell-shaped; cercal plate fused, bilobated at base; bacilliform process straight in apical and medial portion, semicircular at base; surstyli long, simple and medial respect to epandrium ( Fig. 13D ).

Female: body length 3.4-3.6 mm. Differs from male as follows:

Head: Frontal vitta light brown and partially grey pruinose. Ocellar triangle very long, extending up to sixth fr. Frons and vitta broad, the distance between eye margins wider than 0.33x head width. Five to six strong fr, shorter than flagellum, with interstitial setulae, 0.5x length of fr. Upper and lower orb present, and 1 row of frontoorbital setulae; pocl uniformly short, less than 0.5x length of outer and inner vertical setae, the latter longer than in male.

Thorax: Grey in ground colour, brown vitta along full length of acr, dc and ia lines.

Legs: F2 with 1 row of very short ad; 1 row of pv in basal half. T 2 cone-shaped; pubescence on ventral surface absent; with 1 submedial and 1 apical av; 1 submedial and 2 apical pd; 1 apical pv, apical pv absent. F3 slightly broadened at preapex, with 1 row of short av and 2 to 3 subapical av as long as femoral width; ventral and and posteroventral surfaces bare. T 3 with 2 medial ad, apical ad absent; 2 apical and 3 subapical av.

Postabdomen: Cerci normal, short; anal plate as broad as long and covered with setulae; sternite 8 reduced to 1 posterior pair of rounded plates bearing 2 setae and 2 to 3 setulae and 2 smaller anterior plates; post abdominal spiracles 7 and 8 on tergite 6 ( Fig. 13E ). Two pear-shaped, slightly quadrate and grooved spermathecae with partially sclerotized ducts ( Fig. 13F ).

Distribution: NEOTROPICAL: Provinces of San Luis and Mendoza, Argentina.

Discussion: Fannia losgateados does not seem to belong to any of the previously proposed Neotropical species groups of Fannia . It can be easily separated from the remaining species of Fannia of southern South America because of its small size, black colouration and the presence of a row of long and hair-like pv, visible in anterior view on F3.

Figures

FIGURE 13: Fannia losgateados sp. nov. A. Male hind leg, anterior. B. Male sternite 5, ventral. C. Male hypandrium, ventral. D. Male terminalia external structures, ventral. E. Female terminalia external structures, ventral. F. Female terminalia, spermathecae.