Episcomitra pseudoincognita, Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 17-19

publication ID	https://doi.org/10.11646/zootaxa.4983.1.1
publication LSID	lsid:zoobank.org:pub:6A4778D6-195A-4AB1-AA1E-7D8000185B28
DOI	https://doi.org/10.5281/zenodo.5044074
persistent identifier	https://treatment.plazi.org/id/A82A87E9-8A06-385F-FF4D-FE61FF22FDD9
treatment provided by	Plazi (2021-06-10 00:51:57, last updated 2024-11-24 21:02:59)
scientific name	Episcomitra pseudoincognita
status	sp. nov.

Treatment

Episcomitra pseudoincognita View in CoL nov. sp.

Figs 4F, G View FIGURE 4 1 –G View FIGURE 1 2 View FIGURE 2 , H 1 –H View FIGURE 1 2 View FIGURE 2 , I 1 –I View FIGURE 1 2 View FIGURE 2 , 7L View FIGURE 7 1 –L View FIGURE 1 2 View FIGURE 2

[ Mitra ] incognita Bast. —Hoernes 1880: 125 [non de Basterot, 1825].

Mitra incognita Bast. View in CoL — Hoernes & Auinger 1880: 79, pl. 9, figs 3–5 [non de Basterot, 1825].

Mitra Bouéi View in CoL nov. form,— Hoernes & Auinger 1880 (pars): 79, pl. 9, figs 7a–b [non Hoernes & Auinger, 1880].

[ Mitra View in CoL ] incognita Bast. View in CoL — Rzehak 1894: 256 [non de Basterot, 1825].

Mitra Bouei R. Hoern. View in CoL i Auinger— Friedberg 1911: 14, pl. 1, fig. 7 [non Hoernes & Auinger, 1880].

M [itraria]. (M [itraria].) bouéi View in CoL (R. Hörn, et Au.)— Sieber 1958a: 154.

? Mitra cf. incognita View in CoL clavatularis Grat.— Csepreghy-Meznerics 1969: 93, pl. 4, fig. 6 [non Grateloup, 1846].

? Mitra incognita clavatularis Grat. — Csepreghy-Meznerics 1972: 31, pl. 14, fig. 13 [non Grateloup, 1846].

Type material. Holotype: NHMW 1854 View Materials /0035/0100e, SL: 24.3 mm, MD: 8.3 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 4a–b), figs 4G 1 –G 2 . Paratypes: NHMW 1854 View Materials /0035/0100f, SL: 18.4 mm , MD: 7.0 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 3a–b), figs 4H 1 –H 2 ; NHMW 1854 View Materials /0035/0100g, SL: 17.4 mm , MD: 6.7 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 5a–c), figs 4I 1 –I 2 ; NHMW 2020 View Materials /0122/0001, SL: 22.2 mm, 7.5 MD: mm, Lăpugiu de Sus ( Romania), figs 7L 1 –L 2 ; NHMW 1865 View Materials /0015/0014, SL: 21.1 mm, 7.6 , MD: mm, Jerutek at Lysice ( Czech Republic), illustrated in Hoernes & Auinger (1880, pl. 9, figs 7a–b), fig. 4F.

Additional material. NHMW 202 View Materials /0122/0002, 21 specimens, Lăpugiu de Sus ( Romania) .

Type locality. Lăpugiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, early/middle Badenian (Langhian).

Etymology. A combination of Latin pseudo (= wrong) and incognita referring to the misidentification of Mitra incognita de Basterot, 1825 .

Diagnosis. Episcomitra species of medium sized, moderately slender to moderately broad shell with acute conical spire, weakly convex whorls, moderately constricted last whorl, and delicate spiral sculpture of flattish spiral cords.

Description. Shell medium sized, moderately slender fusiform with impressed suture. Protoconch unknown. Teleoconch of eight whorls. Early teleoconch whorls straight sided, conical with five broad, convex spiral cords separated by narrow spiral grooves. Later teleoconch whorls weakly convex, with periphery at abapical suture. Up to ten spiral cords on penultimate whorl developing by bifurcation of primary spiral cords. Spiral sculpture weak or subobsolete on late spire whorls and last whorl in most specimens, rarely persisting throughout. Last whorl weakly convex, with periphery slightly below adapical tip of aperture; moderately constricted with distinct basal concavity. Numerous flattish spiral cords on base and fasciole. Aperture elongate ovoid, moderately wide. Columellar callus thin, narrow, indistinct. Columella with four oblique columellar folds, abapical one strongly reduced. Outer lip thin. Siphonal canal moderately long, wide, slightly bent to the left with deeply incised siphonal notch. Colour pattern in UV light consisting of narrow spiral stripes, disintegrating into long dashes on base and fasciole.

Shell measurements and ratios. SL = 18.4–24.3 mm, MD: 7.0– 8.3 mm, AA = 32–37°, SL/MD: 2.7–3.0, AL/ AW: 4.5–4.7, AH/S: 2.3–2.5.

Discussion. This small species was identified by Hoernes & Auinger (1880) as Mitra incognita de Basterot, 1825 . This identification was already doubted by Bellardi (1887a: 58), without providing an alternative name. We agree with Bellardi (1887a). Episcomitra incognita , from the Burdigalian of the north-eastern Atlantic, has a comparable outline, but is much larger (SL: 36 mm), more solid, and its spiral sculpture is even more delicate (see de Basterot 1825: 45, pl. 4, fig. 5, Peyrot 1928: 99, pl. 9, figs 15–16, 29–30). A specimen from the Aquitanian of France, described by Lozouet et al. (2001) as Mitra incognita , represents a stocky shell with a broad conical spire and therefore is strongly dissimilar to the Paratethyan species. Some Proto-Mediterranean species, originally lumped with Mitra incognita by Bellardi (1850), were later described as separate species ( Episcomitra villalverniensis , Episcomitra afficta ). None of these are conspecific with the Paratethyan species: the Pliocene Episcomitra villalverniensis (Bellardi, 1887) (SL: 27 mm) differs in its shouldered whorls and higher last whorl (see Bellardi 1887a: 44, pl. 3, fig. 20; Ferrero-Mortara et al. 1981: 153, pl. 45, fig. 1). The early Miocene Episcomitra afficta (Bellardi, 1887) (SL: 23 mm) is more slender and has more convex whorls (see Bellardi 1887a: 57, pl. 3, fig. 49; Ferrero-Mortara et al. 1981: 156, pl. 47, fig. 3) and the early Miocene Episcomitra subumbilicata ( Bellardi, 1850) has higher and broader subcylindrical spire whorls (see Bellardi 1887a: 46, pl. 3, fig. 25; Ferrero-Mortara et al. 1981: 154, pl. 45, fig. 5).

Palaeoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (middle Miocene): Ukrainian Fore-Carpathian Basin: Dryszczów (Nadrichne) ( Ukraine) ( Friedberg 1911), Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes and Auinger 1880).

References

Basterot, B. de (1825) Memoire Geologique sur les Environs de Bordeaux. Premiere partie comprenant les observations generales sur les mollusques fossiles, et la description particuliere de ceux qu'on rencontre dans ce bassin. Memoires de la Societe d'Histoire Naturelle de Paris, 2, 1 - 100. [https: // www. biodiversitylibrary. org / item / 23524 # page / 12 / mode / 1 up]

Bellardi, L. (1850) Monografia delle mitre fossili del Piemonte. Memorie della Reale Accademia delle Scienze di Torino, 2 / 11, 357 - 390. [https: // www. biodiversitylibrary. org / item / 32672 # page / 441 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 59630

Bellardi, L. (1887 a) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 5; Mitridae. Ermanno Loescher, Torino, 85 pp. [https: // www. biodiversitylibrary. org / item / 45379 # page / 9 / mode / 1 up]

Csepreghy-Meznerics, I. (1969) Nouvelles gasteropodes et lamellibranches pour la faune hongroise des gisements tortoniens- inferieurs de la Montagne de Bukk. Annales Historico-Naturales Musei Nationalis Hungarici Pars Mineralogica et Palaeontologica, 61, 63 - 127.

Csepreghy-Meznerics, I. (1972) La faune Tortonienne-inferieure des gisements tufiques de la Montagne de Bukk: Gastropodes II. Az Egri Muzeum Evkonyve (Annales Musei Agriensis), 8, 26 - 46.

Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 & 5, 1 - 112, pls. 1 - 5 (1911) & 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]

Grateloup, J. P. S. de (1846) Conchyliologie fossile des terrains tertiaires du Bassin de l'Adour, (environs de Dax). Tome Ier. Univalves. Atlas. Th. Lafargue, Bordeaux, pls. 1 - 45 (1840) + pp. i - xx + 1 - 12, pls. 46 - 48. [https: // archive. org / details / conchyliologief 00 gratgoog]

Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405

Lozouet, P., Lesport, J. - F. & Renard, P. (2001) Revision des Gastropoda (Mollusca) du Stratotype de l'Aquitanien (Miocene inf.): site de Saucats ' Lariey', Gironde, France. Cossmannia, Hors-serie 3, 1 - 190.

Mayer-Eymar, C. (1890) Diagnosis specierum novarum ex agris mollassicis seu neogenis, in Museo Turicensi conservatarum. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zurich, 35, 290 - 301. [https: // www. biodiversitylibrary. org / item / 35342 # page / 304 / mode / 1 up]

Peyrot, A. (1928) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 79, 5 - 263. [https: // www. biodiversitylibrary. org / item / 192271 # page / 155 / mode / 1 up]

Rzehak, A. (1894) Uber einige neue Fossilienfundorte im mahrischen Miocan. Verhandlungen des naturforschenden Vereines in Brunn, 33, 252 - 262. [http: // www. zobodat. at / pdf / Verh-naturf-Ver-Bruenn _ 33 _ 0252 - 0262. pdf]

Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 4. A1–A2. Episcomitra leopoldiana nov. sp., holotype, Steinebrunn (Austria), NHMW 1846/0037/0097a. B1–B2. Episcomitra leopoldiana nov. sp., paratype, Steinebrunn (Austria), NHMW 1846/0037/0097b. C1–C2. Episcomitra leopoldiana nov. sp., paratype, Steinebrunn (Austria), NHMW 2020/0131/0001. D1–D2. Episcomitra praenigra (Mayer-Eymar, 1890), lectotype, Lăpugiu de Sus (Romania), NMB Inv. Nr. t3360. E1–E2. Episcomitra praenigra (Mayer-Eymar, 1890), paralectotype, Lăpugiu de Sus (Romania), NMB Inv. Nr. t3361. F. Episcomitra pseudoincognita nov. sp., paratype, Lăpugiu de Sus (Romania), Jerutek at Lysice (Czech Republic), NHMW 1854/0035/0100. G1–G2. Episcomitra pseudoincognita nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0100e. H1–H2. Episcomitra pseudoincognita nov. sp., paratype, Lăpugiu de Sus (Roma- nia), NHMW 1854/0035/0100f. I1–I2. Episcomitra pseudoincognita nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0100g,

FIGURE 7. Colour patterns in UV light. A1–A2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0002. B1–B2. Domiporta amoena nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0001. C1–C2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 1868/0001/0403. D1–D2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0003. E1–E2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0002. F1–F2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0003. G1–G2. Domiporta pulchra nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0001. H1–H2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0004. I1–I2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0001. J1–J2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0002. K1–K2. Domiporta turpis nov. sp., holotype, Baden (Austria), NHMW 1846/0037/0098ª. L1–L2. Episcomitra pseudoincognita nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0122/0001.