Eocenhecabolus kotenkoi Belokobylskij, 2023

Belokobylskij, Sergey A., Simutnik, Serguei A., Vasilenko, Dmitry V. & Perkovsky, Evgeny E., 2023, First record of the parasitoid subfamily Doryctinae (Hymenoptera, Braconidae) in Rovno amber: description of a new genus and species with stigma-like enlargement on the hind wing of the male, Journal of Hymenoptera Research 95, pp. 59-72 : 59

Eocenhecabolus kotenkoi Belokobylskij sp. nov.

Figs 1 , 2

Type material.

Holotype: male, SIZK VT-607, Velyki Telkovichi, Varash District, Rovno amber, late Eocene.

Description.

Male. Body length 1.5 mm; fore wing length 1.3 mm.

Head: Head relatively high, its width about 1.3 times medial length. Occiput weakly concave. Transverse diameter of eye 1.5 times longer than temple (subdorsal view). POL 1.3 times Od, approximately 0.5 times OOL. Eye about 1.2 times as high as broad (lateral view). Malar space 0.3 times height of eye, almost equal to basal width of mandible. Face width 0.9 times height of eye, 1.3 times medial height of face and clypeus combined. Hypoclypeal depression subround, its transverse width 0.9 times distance from edge of depression to eye, 0.4 times width of face.

Antenna: First flagellomere almost 7.0 times longer than its apical width, approximately twice longer than scape. Second segment present only basally, remaining part missing.

Mesosoma: Mesosoma long, its length 1.8 times height. Neck of prothorax relatively short. Pronotal carina absent, dorsal pronotal lobe distinctly convex. Median lobe of mesoscutum convex, distinctly protruding forward, without anterolateral corners. Prescutellar depression relatively long. Subalar depression shallow and mainly smooth. Lateral carinae between propodeum and metapleuron strong and complete.

Wings: Fore wing wide, 2.6 times longer than its maximum width. Pterostigma wedge-shaped, 3.7 times longer than its width. Radial vein (r) arising from basal 0.4 of pterostigma. First (r) and second (3RSa) radial abscissae forming obtuse angle; first abscissa (r) 0.7 times as long as maximum width of pterostigma. Second radial abscissa (3RSa) 3.0 times first abscissa (r), 0.5 times as long as the straight third abscissa (3RSb), 1.3 times longer than the straight first radiomedial vein (2RS). Second radiomedial (submarginal) cell relatively wide and long, 2.7 times longer than its maximum width, 1.8 times longer than the narrow brachial (first subdiscal) cell. Recurrent vein (1 m-cu) 0.75 times as long as first radiomedial vein (2RS), 0.6 times as long as basal vein (1M). Discoidal (first discal) cell rather short, 1.7 times longer than its maximum width. Nervulus (1cu-a) 0.6 times as long as distance between basal (1M) vein and nervulus (1cu-a). Parallel vein (2CUb) weakly curved basally. Brachial (second subdiscal) cell relatively short and narrow. Hind wing almost 4.5 times longer than its maximum width. Stigma-like enlargement 3.5 times longer than maximum width. First abscissa of mediocubital vein (M+CU) almost twice longer than second abscissa (1-M).

Legs: Fore femur about 4.5 times longer than maximum width. Fore tarsus 1.2 times longer than fore tibia. Hind coxa almost 1.5 times longer than its maximum width, 0.8 times as long as propodeum. Hind femur 3.0 times longer than its width. Hind tarsus almost as long as hind tibia. Second segment of hind tarsus 0.4 times as long as basitarsus, weakly longer than fifth segment (without pretarsus).

Metasoma: Length 1.2 times larger than length of head and mesosoma combined. First metasomal tergite 1.4 times longer than distal maximum width, 1.3 times longer than propodeum; apical width of first tergite about 1.6 times its basal width. Second and third tergites combined 1.3 times longer than basal width of second tergite, 0.9 times as long as their maximum width.

Sculpture and pubescence: Temple densely transversely and sinuately striate with additional reticulation laterally. Face weakly transversely striate, smooth medially. Frons and most part of temple perhaps mainly smooth. Propodeum mostly smooth, only sometimes with short and sparse rugae along carinae; areola almost 2.5 times longer than its width; basomedial carina present in basal 0.3 of propodeum. Hind coxa and femur smooth. First metasomal tergite striate medially, weakly rugose sublaterally, almost smooth laterally. Second tergite mainly smooth, finely striate in small basolateral areas. Remaining part of metasoma smooth. Hind tibia with rather dense and short semi-erect setae, its length 0.4-0.6 times maximum width of tibia.

Colour: Body almost entirely brown. Legs mainly reddish brown to pale reddish brown. Fore wing almost entirely faintly evenly infuscate. Pterostigma entirely brown.

Female. Unknown.

Etymology.

This species is named in honour of the well-known Ukrainian braconidologist, Dr Anatoly Grigorievich Kotenko.

Figures

Figure 1. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A habitus, right dorso-lateral view B habitus, left ventro-lateral view C body, lateral view D body, ventro-lateral view E head and antenna, fronto-lateral view F head, dorsal view G propodeum and metasoma, dorsal view H head and antenna, lateral view.

Figure 2. Eocenhecabolus kotenkoi gen. et sp. nov. (male, holotype, Rovno amber, # SIZK VT- 607) A wings B metasoma and hind leg, lateral view.