Enspeletoma oligocaenica Rasnitsyn & Wedmann, 2024

Sun, Chenhui, Rasnitsyn, Alexandr P., Wedmann, Sonja, Zhuang, Jialiang, Shih, Chungkun, Ren, Dong & Gao, 2024, Revisions * new taxa * and venation transformations of the sawfly family Blasticotomidae sensu lato (Hymenoptera: ºenthredinoidea) highlight the evolution of the basal Hymenoptera, Zoological Journal of the Linnean Society 202 (1), pp. 1-19 : 3-6

Enspeletoma oligocaenica Rasnitsyn & Wedmann gen. et sp. nov.

( Fig. 2A–C )

http://zoobank.org/urn:lsid:zoobank.org:act:B0B4637E-607D-4B87-A3C5-4F842B05C035

Diagnosis: As for genus (see above).

Etymology: The species is named after the Oligocene* the geological epoch when it was fossilized.

Material: Holotype * coll. no. NHMMZ PE 1995/9108-LS * only one plate. Dorsoventrally embedded* parts of thorax* large parts of left forewing and basal parts of right forewing preserved ( Fig. 2A ); forewing venation can be reconstructed with combined information from both wings ( Fig. 2B * C).

Locality and horizon: Enspel * Westerwald * Germany; Digging site 5* Local horizon 16 ( Wedmann 2000); Upper Oligocene* 24.9–24.5 Mya; Mammal Palaeogene zone MP 28 (Storch et al. 1996* Mertz et al. 2007).

Description: Holotype * NHMMZ PE 1995/9108-LS. Sex unknown. Thorax 3.7 mm long as preserved. Pterostigma dark* very large* ~ 1.1 mm long* 0.6 mm wide. Costal space narrow as preserved* with no traces of Sc visible. Vein 1-Rs distinct* short. 1-M* 1-Cu* and 2-M & RS+M (upper side of cell 1mcu) of almost equal lengths. Rs between 2r-rs and 2r-m much shorter than 2r-rs and slightly longer than 1m-cu. Vein 2r-m about as long as vein 2r-rs* and 2r-m equidistant from 1m-cu and 2m-cu. Cell 2mcu relatively short* about as long as cell 1r+2rm. Cell 1r+2rm as long as cell 3rm. Distal part of cell 3rm apparently very high (higher than distal part of cell 2mcu). 1m-cu about half as long as 3-Cu and 4-Cu. 2-Cu ~0.6 times as long as 1-Cu and ~1.4 times as long as1m-cu.

Figures

Figure 2. Cenozoic extinct Blasticotominae.A–C* Enspeletoma oligocaenica. Holotype* NHMMZ PE 1995/9108-LS* sex unknown* Upper Oligocene of Enspel* Germany.A* dorsal view.B* C* line drawings of forewing* reconstructed from both preserved wings* with veins labelled (B) and with cells labelled (C). D* E* Paremphytus ostentus. Holotype No. 2042* Museum of Comparative Zoology* Harvard University* Cambridge* MA* USA; female* uppermost Eocene* Colorado* USA. D* dorsal view.E* line drawing (Zhelochovtzev and Rasnitsyn 1972). Scale bars:1 mm.