Elaphoglossum sect. Setosa

Kessler, Michael, Moran, Robbin C., Mickel, John T., Matos, Fernando B. & Smith, Alan R., 2018, Prodromus of a fern flora for Bolivia. XXXV. Dryopteridaceae, Phytotaxa 353 (1), pp. 448-450 : 448-450

Key to the Bolivian species of Elaphoglossum sect. Setosa

1 Scales on the abaxial blade surfaces flat (use magnification!); sterile leaves and rhizomes sometimes with white patches or spots; blades simple or pedate (sect. Undulata sensu Mickel & Atehortúa, 1980 ) .................................................................................... 2

– Scales on the abaxial blade surfaces between costae and margins subulate; sterile leaves and rhizomes without white patches or spots; blades simple (sect. Setosa sensu Mickel & Atehortúa, 1980 ) ............................................................................................ 11

2 Blades pedate ................................................................................................................................................................ E. cardenasii

– Blades simple .................................................................................................................................................................................... 3

3 Petioles dark brown to castaneous; bases of sterile blades short-decurrent with concave sides ................................. E. papillosum

– Petioles stramineous to light brown; bases of sterile blades various ................................................................................................ 4

4 Bases of sterile blades cordate ............................................................................................................................... E. cardioglossum

– Bases of sterile blades truncate, rounded, cuneate, or long-decurrent ............................................................................................. 5

5 Sterile blades proliferous at the apices ............................................................................................................................. E. wardiae

– Sterile blades not proliferous ............................................................................................................................................................ 6

6 Petioles <1/20 the sterile blade length; white patches on abaxial blades present, restricted to the leaf tissue between veins ............ ........................................................................................................................................................................................ E. zebrinum

– Petioles 1/4–1/2 the sterile blade length; white patches on abaxial blades absent or, if present, extending across veins ................ 7

7 Rhizomes long-creeping (but leaves may be clustered at the rhizome apices) ................................................................................ 8

– Rhizomes short- to very short-creeping ........................................................................................................................................... 9

8 Sterile blades 12 – 20 × 3–5 cm, firm to subcoriaceous, clustered at rhizome apices; plants terrestrial or saxicolous ... E. herpestes

– Sterile blades 10 – 14 × 1–2 cm, membranaceous, ca. 1 cm apart along the rhizomes; plants usually epiphytic ...... E. squarrosum

9 Scales on the abaxial blade surfaces ca. 0.3 mm long, rounded to substellate; petiolar scales spreading, usually reflexed; intersporangial scales lacking ................................................................................................................................... E. beaurepairei

– Scales on the abaxial blade surfaces ca. 1 mm long, lanceolate to linear-lanceolate; petiolar scales appressed to spreading; intersporangial scales present ......................................................................................................................................................... 10

10 Bases of sterile blades long-decurrent ............................................................................................................................ E. delasotae

– Bases of sterile blades cuneate to rounded ...................................................................................................................... E. notatum

11 Apices of sterile blades rounded to acute ....................................................................................................................................... 12

– Apices of sterile blades cuspidate, acuminate, or gradually attenuate ........................................................................................... 13

12 Fertile blades narrowly oblanceolate, often inrolled; adaxial blade scales orangish; fertile leaves about the length of the sterile ones; 200 – 1500 m ....................................................................................................................................................... E. horridulum

– Fertile blades round to ovate, flat or conduplicate; adaxial blade scales black to orangish; fertile leaves longer than the sterile ones; (1300 –) 2000 – 3200 m ................................................................................................................................................ E. piloselloides

13 Rhizomes long-creeping; rhizome scales widely patent .................................................................................... E. heteromorphum

– Rhizomes short- to very short-creeping; rhizome scales appressed to slightly patent ................................................................... 14

14 Sterile blades 2 – 7 cm long, apically proliferous .............................................................................................. E. didymoglossoides

– Sterile blades 5–60 cm long, not proliferous .................................................................................................................................. 15

15 Sterile blades 5–15 cm long ........................................................................................................................................................... 16

– Sterile blades 30–60 cm long ......................................................................................................................................................... 19

16 Blades narrow-lanceolate; petioles <1/4 the sterile leaf length ....................................................................................... E. rosettum

– Blades oblong to ovate; petioles>1/2 the sterile leaf length .......................................................................................................... 17

17 Sterile blades ovate, apically acuminate, basally cordate .............................................................................................. E. solomonii

– Sterile blades oblong to lanceolate, apically cuspidate, basally truncate to subcordate ................................................................. 18

18 Sterile blade bases basally cuneate to rounded; petioles and blades glabrous to sparsely scaly ...................................... E. lindenii

– Sterile blade bases basally truncate to subcordate; petioles and blades conspicuously scaly .......................................... E. pilosius

19 Petioles with subulate scales and small, glandular hairs 0.1–0.3 mm long .................................................................................... 20

– Petioles with only long subulate scales, lacking glandular hairs .................................................................................................... 21

20 Sterile blades 2.5–4 cm wide ........................................................................................................................................ E. catenatum

– Sterile blades 1.5–2.3 cm wide ...................................................................................................................................... E. haynaldii

21 Petioles 1/20–1/10 the sterile leaf length; bases of sterile blades long-decurrent ................................................. E. poeppigianum

– Petioles ca. 1/3 the sterile leaf length; bases of sterile blades rounded to cuneate ......................................................................... 22

22 Blades linear-lanceolate, basally rounded to truncate ................................................................................................... E. setigerum

– Blades lanceolate to elliptic, basally narrowly cuneate .................................................................................................................. 23

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