Draconarius postremus, Wang & Jäger, 2008

Wang, Xin-Ping & Jäger, Peter, 2008, First record of the subfamily Coelotinae in Laos, with review of Coelotinae embolus morphology and description of seven new species from Laos and Vietnam (Araneae, Amaurobiidae), Journal of Natural History 42 (35 - 36), pp. 2277-2304 : 2294-2297

Draconarius postremus sp. nov.

( Figures 44–57 , 75, 76 )

Type material

Holotype female and one paratype male from Muang Sing , Nam Det, Luang Nam Tha Province, Laos, 821–1097 m, 21 ° 10.1939N 101 ° 14.4459E to 21 ° 09.9839N 101 ° 14.7429E, secondary forest, along path, soil and vegetation, hand, sweepnet, sieving, P. Jäger & V. Vedel leg., 6 November 2004, deposited in SMF.

Etymology

The specific name is derived from the Latin adjective ‘‘postremus’’, meaning ‘‘most posterior’’, and referring to the posterior extension of the atrium; adjective.

Diagnosis

Females of this new species can be easily recognized by the tiny epigynal teeth situated laterally of the atrium close to the epigastric furrow, the small, posterior extending atrium, and the strongly convoluted, laterally extending spermathecae ( Figures 44, 45 , 49, 50 ), and males by the minute patellar apophysis, the short conductor, the simple, broad and not spoon-shaped median apophysis, and the embolic base arising retrolaterally ( Figures 46–48 ).

Description

Female holotype. Small spider, total length 4.25 mm ( Figures 51–53 ). Dorsal shield of prosoma 2.10 long, 1.35 wide; opisthosoma 2.15 long, 1.30 wide. AME smallest, approximately half the size of other eyes (AME 0.04, ALE 0.09, PME 0.08, PLE 0.09); AME, ALE, PLE close together, separated by slightly less than AME diameter, other eyes are relatively widely separated by slightly less than PME diameter (AME–AME 0.03, AME–ALE 0.03, ALE–PLE 0.04, PME–PME 0.07, PME–PLE 0.06, AME–PME 0.06) ( Figure 56 ). Chelicera with three promarginal and two retromarginal teeth. Epigynal teeth minute, indistinct, situated posteriorly close to epigastric furrow and widely separated by more than atrial width; atrium small, situated and extending posteriorly; copulatory ducts originating posteriorly and extending anteriorly, indistinct from dorsal view; spermathecae large, close together, looping and extending laterally; spermathecal heads small, situated distally of spermathecal lateral extension ( Figures 44, 45 , 49, 50 ).

Male paratype. Small spider, total length 4.00 mm ( Figures 54, 55 ). Dorsal shield of prosoma 1.95 long, 1.25 wide; opisthosoma 2.10 long, 1.30 wide. Eyes are similar to female eyes, with AME smallest, approximately half the size of other eyes; AME, ALE, PLE separated by slightly less than AME diameter, other eyes are separated by slightly less than PME diameter ( Figure 57 ). Chelicera with three promarginal and two retromarginal teeth. Palp with a tiny patellar apophysis; retrolateral tibal apophysis slightly longer than half of tibial length; lateral tibial apophysis broad; cymbial furrow long, extending more than half of cymbial length; conductor short, strongly concave, with broad lamella and large dorsal apophysis, its sheath-like part with a subdistal indentation; median apophysis broad, not spoon-shaped; embolus (type III) long, retrolateral in origin, arising in a 5 o’clock position and extending proximally beyond tibia/tarsus junction, running two-thirds of an oval, extending and coiling distally beyond distal bulb ( Figures 46–48 ).

Relationships

The only Coelotinae that might be related to D. postremus sp. nov. is D. rufulus (Wang et al., 1990) described from Zhejiang and Anhui Provinces in southeastern China. Both species share the small, posteriorly extending atrium and the large, convoluting spermathecal tubes, but D. rufulus has no patellar apophysis, exhibits a spoon-shaped median apophysis and the strongly modified conductor and embolus. From the latter characters it is unlikely that they form a sister group relationship.

Distribution

Laos (Luang Nam Tha Province: Nam Det) ( Figures 75–76 ).

Figures

Figures 44–48. Draconarius postremus sp. nov., female holotype (44–45) and male paratype (46–48) from Muang Sing, Nam Det, Luang Nam Tha Province, Laos. 44. Epigynum, ventral view. 45. Vulva, dorsal view. 46. Palp, prolateral view. 47. Palp, ventral view. 48. Palp, retrolateral view.

Figures 49, 50. Draconarius postremus sp. nov., female holotype from Muang Sing, Nam Det, Luang Nam Tha Province, Laos. 49. Epigynum, ventral view. 50. Vulva, dorsal view.

Figures 51–53. Draconarius postremus sp. nov., female holotype from Muang Sing, Nam Det, Luang Nam Tha Province, Laos. 51. Habitus, dorsal. 52. Habitus, lateral. 53. Habitus, ventral view.

Figures 54–57. Draconarius postremus sp. nov., female holotype (56) and male paratype (54, 55, 57) from Muang Sing, Nam Det, Luang Nam Tha Province, Laos. 54. Habitus, dorsal view. 55. Habitus, ventral view. 56, 57. Eyes, dorsal view.

Figures 75, 76. 75. Records of Draconarius bounnami sp. nov., D. latidens sp. nov., D. postremus sp. nov., D. songi sp. nov., D. tabularis sp. nov. from Luang Nam Tha Province, Laos and D. hanoiensis sp. nov. from Hanoi, Vietnam and D. huongsonensis sp. nov. from Huong Son, Vietnam. 76. Records of five species from Luang Nam Tha Province, Laos, enlarged.