Dissochaeta intermedia Blume, Flora 14: 493. 1831.

Kartonegoro, Abdulrokhman, Veldkamp, Jan Frits, Hovenkamp, Peter & Welzen, Peter van, 2018, A revision of Dissochaeta (Melastomataceae, Dissochaeteae), PhytoKeys 107, pp. 1-178 : 1

28. Dissochaeta intermedia Blume, Flora 14: 493. 1831. Map 17

Dissochaeta monticola Blume, Flora 14: 494. 1831. Type: Indonesia. West Java: Kuripan, C.L. Blume s.n. (lectotype, designated here: L [L0537300]!; isolectotypes: L [L0537301, L0537302]!).

Type.

Indonesia. West Java: Preanger Regentsch., Mount Pangrango, Gegerbentang, C.L. Blume 539 (lectotype, designated here: L [L0537299]!; isolectotypes: K [K000859493, K000859494]!, L [L0537296, L0537297, L0537298]!, P [P05283548, image seen]!).

Description.

Climbing up to 20 m in height. Branchlets terete, 4-7 mm diameter, glabrescent to densely stellate-furfuraceous; nodes swollen, with interpetiolar ridge or line; internodes 8-9 cm long. Leaves: petioles flattened, 10-20 mm long, glabrescent to densely stellate-furfuraceous; blades ovate to ovate-elliptic, 7-16 × 5-8.5 cm, membranous, base rounded, margin entire, apex acuminate, tip 0.5-2 cm long; nervation with 1 pair of lateral nerves and 1 pair of intramarginal nerves; adaxially glabrous, abaxially densely brown stellate-tomentose. Inflorescences terminal and in the upper leaf axils, up to 32 cm long, many-flowered; main axis angular, densely stellate-furfuraceous; primary axes up to 25-30 cm long with 4-6 nodes, secondary axes 2-7 cm long with 2 or 3 nodes, tertiary axes 1-3 cm long with 1 or 2 nodes, quartenary axis up to 0.8 cm long when developed with 1 node; bracts minute, inconspicuous; bracteoles linear to ovate-oblong, 1-2 mm long, densely stellate-furfuraceous; pedicels densely stellate-furfuraceous, 4-7 mm long in central flowers, 2-4 mm long in lateral flowers. Hypanthium campanulate, slightly angular, 2-5 × 1-3 mm, with four distinct ridges, stellate-puberulous to densely stellate-tomentose, somewhat pointing sideways from pedicels; calyx lobes truncate with undulate tips, 0.5-1 mm long, widened, stellate-furfuraceous; petal bud conical 3-7 mm long; mature petals ovate or oblong, 6-10 × 3-5 mm, base clawed, apex obtuse, glabrous or inside at base with appressed hairs, pink or violet. Stamens 4, equal, alternipetalous, filaments flat, 4-6 mm long, straight, apex bent, anthers linear-lanceolate, sickle- or S-shaped or falcate, thecae 5-6 mm long, yellow, pedoconnective ca. 1 mm long, basal crests minute, triangular, ca. 1 mm long, narrow with acute apex, lateral appendages paired, filiform, 3-4 mm long, tan. Ovary half or ⅔ of hypanthium in length, apex pubescent; style erect, 8-12 mm long, curved at apex, glabrous, purple; stigma minute; extra-ovarial chambers 4, alternipetalous, extending to the base of the ovary. Fruits ovoid, 5-6 × 3-5 mm, glabrous to sparsely stellate-furfuraceous, rarely with distinct vertical ridges, green, often pointing sideways from the pedicels; calyx lobe remnants persistent. Seeds ca. 0.5 mm long.

Distribution.

Java (West).

Ecology and habitat.

Primary montane forest or in edge of forest in open, 1200-2000 m elevation.

Vernacular names.

harendong cai, harendong areuy, harendong kowe (Sunda).

Note.

Dissochaeta intermedia resembles D. leprosa in the number and shape of the stamens, but is different in having a smaller hypanthium and truncate calyx lobes with undulate tips. Together with D. leprosa , this species is common at high elevations up to 2000 m in Java.

Specimens examined.

INDONESIA. West Java: Bogor, Puncak Pass, 1300 m, 3 Mar 1927, J.G.B. Beumée A390 (BO); Ibid., Cisarua, 1200 m, 19 Feb 1950, C.G.G.J. van Steenis 12735 (L); Mt. Salak, 1200 m, H. Raap 158 (L); Kuripan, C.L. Blume s.n. (L); Megamendung, H. Kuhl & J.C. van Hasselt s.n. (L); Ibid., F.W. Junghuhn s.n. (U); Mt.Halimun, Nirmala, 1250 m, 9 Jun 1980, M.M.J. van Balgooy & H. Wiriadinata 2889 (BO, L); Ibid., Mt. Bintang Gading, 19 May 2002, W.S. Hoover et al. 5502 (BO); Cianjur, Mt.Gede, Cibodas, J.G. Boerlage s.n. (L); Ibid., 1400 m, 22 Aug 1879, Arsin 19556 (BO); Ibid., 9 Jun 1906, A.A. Pulle 4073 (U); Ibid., 19 Jul 1913, S.H. Koorders 42153β (BO); Ibid., 6 May 1914, J.A. Lörzing 1493 (BO); Ibid., 14 Feb 1915, H.N. Ridley s.n. (K); Ibid., Gunung Putri, 6 Nov 1896, S.H. Koorders 25927β (BO); Ibid., Sindanglaya, 1400 m, 6 Jun 1917, C.A. Backer 22286 (BO); Ibid., J.G. Hallier 626 (BO); Ibid., 25 Aug 1893, J.G. Hallier 432 (BO); Ibid., 1915, Sapiin 57 (BO); Ibid., 15 Jul 1908, T. Valeton s.n. (BO); Ibid., Mt. Pangrango, P. de Monchy s.n. (BO); Ibid., Geger Bentang, C.L. Blume 539 (K, L, P); Bandung, Mt. Tangkuban Perahu, 1700 m, 4 Apr 1912, C.A. Backer 2387 (BO); Ibid., 1600 m, 5 Mar 1912, C.A. Backer 2415 (BO); Ibid., Jul 1915, W.M. Docters van Leeuwen 2301a (BO); Ibid., 24 Jul 1927, W.M. Docters van Leeuwen 11423 (BO); Ibid., 2000 m, 21 Nov 1952, W. Meijer 1363 (BO); Ibid., 28 May 1908, H.H. Zeijlstra 19 (BO); Ibid., 6 Feb 1927, C.A. Wisse 1188 (BO); Mt. Patuha, Telaga Patengan, 1700 m, 28 Mar 1914, C.A. Backer 12787 (BO); Mt. Burangrang, Situ Lembang, 1600 m, 24 Jul 1920, R.C. Bakhuizen van den Brink 4557 (BO, K, L); Ibid., 30 Dec 1956, S. Reksodihardjo 6 (BO); Mt. Malabar, 27 Jun 1871, R.H.C.C. Scheffer s.n. (BO); Mt. Mandalagiri, 1570 m, 29 Mar 1920, H.J. Lam 230 (BO).

Figures

Map 17. Distribution of D. hirsutoidea (●), D. horrida (■) and D. intermedia (▲).