Dendarus leonhardi Schuster, 1940

Nabozhenko, Maxim V. & Poggi, Roberto, 2022, Contribution to the knowledge of the genus Dendarus Dejean, 1821 (Coleoptera: Tenebrionidae: Blaptinae: Dendarini) from Iran, Turkmenistan and some adjacent territories, Zootaxa 5155 (1), pp. 105-123 : 110

Dendarus leonhardi Schuster, 1940

( Figs 3 , 9C , 10C , 11C , 12C , 13C )

Schuster, 1940: 861; Kühnelt, 1957: 77; Medvedev, 1968: 97, 98, 106; Medvedev & Nepesova, 1985: 122; Medvedev & Nepesova, 1990a: 25; Medvedev & Nepesova, 1990b: 886.

Type material. Lectotype, ♂ ( NMB), designated here: “TYPUS”, “ leonhardi ”, “ Persia Astrabad 5.99 Coll. Hauser.”, “Sammlung Adr. Schuster”. The following label was added to the specimen: “ Lectotypus Dendarus leonhardi Schuster, 1940 des. M. Nabozhenko 2022”.

Material. Turkmenistan. 1♂, 1♀ ( ZIN): Gasan-Kuli Natural Reserve [coast of the Caspian Sea], 30.i.1975 (leg. Yeskov) .

Diagnosis. Male. Body dull, robust. Head and pronotum with coarse and dense uniform puncturation of round punctures (puncture diameter slightly longer or subequal to interpuncture distance). Head ventrally with simple (not impressed) longitudinal furrow beyond prementum. Pronotum weakly convex. Lateral margins of pronotum not emarginated at basal quarter; anterior angles not protruding; base deeply bisinuate, posterior angles acute, protruding, but not extending beyond base of pronotum. Prosternal process comparatively wide, widely rounded at apex. Elytra widest at middle. Strial punctures large, sparse (19–24 punctures in one row), round to weakly elongate, located each in wide deep impression (but impressions not as deep and large as in D. crenulatus ), much larger than interstrial ones. Interstriae with fine and sparse puncturation. Protibiae moderately wide, with deep notch on inner side and right-angled tooth before indentation; protibia widest at level of tooth, where it is slightly wider than at apical level. Mesotibiae straight or very weakly bending inwards. Mesotarsomeres 1–4 very weakly widened, tarsomeres 1 and 3 with reduced hair brush on sole, tarsomere 2 with full hair brush.

Female. Body more robust, protibiae weakly bent. Pronotum shinier than head and elytra and with finer and sparser puncturation than on head. First three strial rows with strongly elongate narrow punctures in form of sparse dotted lines, other striae with slightly elongate or round punctures.

Distribution. NE Iran, SW Turkmenistan.

Figures

FIGURE 3. Dendarus leonhardi. A = lectotype, male, dorsally; B = ditto, ventrally; C = labels of the lectotype; D = lectotype, male, laterally; E = male (non-type), dorsally; F = female (non-type), dorsally.

FIGURE 9. Dendarus spp., male protibia. A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. simplex; E = D. matthewsi sp. n.; F = D. transcaspicus; G = D. vagabundus.

FIGURE 10. Dendarus spp., male tergite VIII. A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = D. transcaspicus; F = D. vagabundus.

FIGURE 11. Dendarus spp., male inner sternite VIII.A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = D. transcaspicus; F = D. vagabundus. sVIII—inner sternite VIII, tVIII—inner tergite VIII, tIX—inner tergite IX.

FIGURE 12. Dendarus spp., spiculum gastrale.A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = D. transcaspicus; F = D. vagabundus.

FIGURE 13. Dendarus spp., aedeagus, ventrally. A = D. armeniacus; B = D. crenulatus; C = D. leonhardi; D = D. matthewsi sp. n.; E = the same, turned forward; F = D. transcaspicus; G = D. vagabundus.