Coprophilus (s. str.) maximus Gildenkov, 2022

Gildenkov, M. Yu., 2022, New species of the genus Coprophilus Latreille, 1829 from North India (Coleoptera: Staphylinidae: Oxytelinae), Russian Entomological Journal 31 (2), pp. 140-143 : 141-143

Coprophilus (s. str.) maximus Gildenkov , sp. n.

Fig. 1 .

MATERIAL. Holotype, ♀, North India , Uttaranchal : with labels “N INDIA: Uttaranchal state, 30 km N Bageshwar, Khati vill env., 2100–2300m, 27–30.VI.2003, Kejval & Tryzna” (cMSch).

DESCRIPTION (holotype). Length 8.8 mm. Body slightly convex, the integument integument slightly shining. Colouration brown, with a reddish tint; antennae black-brown ( Fig. 1 ). Head slightly transverse, widest across eyes. Ratio of its length (from base to anterior margin of clypeus) to maximum width (mm) about 1.07:1.39. Eyes moderately large, slightly convex ( Fig. 1 ). Forehead finely and sparsely punctate. ( Fig. 1 ); vertex with noticeably larger and denser punctation ( Fig. 1 ). Puncture diameter on the vertex is about 1.5 times more than diameter of the eye facet and diameter of points on the forehead, distances between punctures noticeably more than their diameter, interspaces smooth, slightly shining ( Fig. 1 ). Neck large, coarsely and densely punctate. Puncture diameter is about 3 times more than diameter of eye facet, distances between punctures noticeably smaller than their diameter, interspaces smooth, slightly shining. Antennae rather long ( Fig. 1 ). Antennal segment 1 elongate, conical, more than twice longer than its maximum width; segment 2 elongate, conical, about twice as long as its maximum broad, much shorter and narrower than segment 1; segment 3 conical, more than twice longer than its maximum broad; segments 4–6 cylindrical, slightly broaded to apex, only slightly longer than their maximum broad; segments 7–10 conical, much more massive than segments 4–6, slightly transverse; segment 11 pointed apically, about 1.5 times as long as broad. Last 5 segments form loose club ( Fig. 1 ).

Pronotum ( Fig. 1 ) slightly convex, maximum broad after about 2/3 of length, measured from the base. Ratio of pronotum length to its maximum broad (mm) about 1.63:1.60. Front angles rounded, lateral margin serrated ( Fig. 1 ). Pronotal disc with deep depressions: 2 paired oval depressions at base, on either side of midline; unpaired broad rhomboid depression at disc centre and 2 broad, deep, triangular depressions near lateral margins ( Fig. 1 ). Surface of pronotum with delicate, fine and sparsely punctation. Punctation irregular, the diameter of most punctures approximately 1.5 times more than diameter of eye facet, some punctures approximately equal to the diameter of eye facet; the distances between punctures are much longer than their diameter, interspaces smooth, slightly shining ( Fig. 1 ). Punctures at the base of pronotum and along the lateral margins are larger and denser; their diameter about 3 times more than diameter of eye facet, distance between punctures significantly smaller than their diameter. Punctation most dense at basal angles and at bottom of depressions near lateral margins, interspaces between these punctures much smaller than their diameter.

Elytra convex, ratio of length of elytra to their combined width (mm) about 2.20:2.46. Six rather shallow grooves are clearly visible on the surface of each elytron ( Fig. 1 ). Surface of elytra delicate, finely and sparsely punctate. Punctation irregular, diameter of most punctures approximately equal to diameter of eye facet, some punctures are 1.5 times more than diameter of eye facet, distances between punctures are much greater than their diameter, interspaces smooth, slightly shining ( Fig. 1 ). Elytra finely bordered.

Abdomen with delicate, fine and sparsely punctation and delicately shagreened ( Fig. 1 ).

Male. Unknown.

COMPARATIVE NOTES. The new species is most similar in color and general habit to C. (s. str.) formosanus ( Figs 1– 3 ). Well differs from it by significantly larger body sizes. Basic dimensions C. maximus : body length 8.8 mm; length of pronotum 1.63 mm; maximum width of pronotum 1.60 mm, length of elytra 2.20 mm; maximum width of elytra 2.46 mm. Basic dimensions C. formosanus (♀, Paratype, Fig. 2 ): body length 8.0 mm; length of pronotum 1.23 mm; maximum width of pronotum 1.52 mm, length of elytra 1.96 mm; maximum width of elytra 2.15 mm. Basic dimensions C. formosanus (♂, Holotype, Fig. 3 ): body length 7, 7 mm; length of pronotum 1.24 mm; maximum width of pronotum 1.54 mm, length of elytra 1.86 mm; maximum width of elytra 2.00 mm. From a rather large species of the genus and widely known in Europe C. striatulus , also well differs by significantly larger sizes: Basic dimensions C. striatulus (♀, Russia, Smolensk. Fig. 4 ) body length 7.0 mm; length of pronotum 1.13 mm; maximum width of pronotum 1.32 mm, length of elytra 1.72 mm; maximum width of elytra 1.62 mm. The new species well differs from C. formosanus by more delicate and fine punctation of head, pronotum, and elytra, and by less deep striae on the elytra ( Figs 1–3 ). The new species well differs from C. formosanus and C. striatulus by the wide, only slightly longer than maximum broad unpaired rhomboid impression in center of pronotal disc; in C. formosanus and C. striatulus impression is much narrower and more elongated, more than 2 times longer as broad ( Figs 1–4 ).

DISTRIBUTION. North India, Uttaranchal state.

ETYMOLOGY. From Latin “maximum” (greatest); the name is given due to the very large size.

Acknowledgements. The author wishes to thank all colleagues for making material available for study: Michael

Schülke (Berlin), Yasutoshi Shibata (Machida, Tokyo),Alexey Shavrin ( Daugavpils University Institute of Systematic Biology ), Tadashi Ishikawa ( TULE). I also thank Kirill Makarov for taking the photographs (Moscow Pedagogical State University, Russia) .

Figures

Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм.