Colocistis chemsaki Kimsey & Wasbauer, 2013

Kimsey, Lynn S. & Wasbauer, Marius S., 2013, Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae), Journal of Hymenoptera Research 33, pp. 1-24 : 7-8

Colocistis chemsaki Kimsey & Wasbauer sp. n. Figures 3 , 13, 16 , 37 , 43 , 51

Holotype male.

Mexico: Baja California Sur, La Paz, 21/ix.1967, J. A. Chemsak, on flowers during day (EMEC).

Paratypes.

1 male, 15 mi w La Paz, Michelbacher and Ross; 1 male, Santo ( “San”) Domingo, Michelbacher and Ross (BME, CAS).

Diagnosis.

This is one of three species, including Colocistis eremi and Colocistis brevis , with only two forewing submarginal cells. Unlike Colocistis eremi the gular carina is simple, and unlike Colocistis brevis the aedeagus is not decurved. This species shares several features with Colocistis pilosa . The ocelli are small, about the size of the pedicel, whereas they are much larger than the pedicel in other Colocistis , and the eyes closely follow the head margins as opposed to bulging laterally. Unique features include the antenna darker than the rest of the body, whereas they are the same color in the other species, and the propodeum in Colocistis chemsaki has two large elevated, ovoid areas on either side of the midline of the dorsal surface.

Description.

Male.

Body length. 7-8 mm.

Head ( Fig. 3 ). Compound eyes slightly enlarged, barely extending laterally beyond temples, inner margins somewhat emarginated; ocelli small, diameter subequal to pedicel length; of lateral ocellus at least 0.7 × OOL, removed from vertex by 0.5 or less an ocellar diameter in frontal view; impressed line present between lateral ocelli posteriorly; clypeus nearly flat, not visible above mandibles in lateral view, without projecting tubercle, apex not margined by strong teeth; antennal socket rim only slightly thickened ventrolaterally; LID 0.9 × UID; flagellomere I length 2.3 × breadth; flagellomere II length 2.8 × breadth; flagellomere IX length 4.2 × breadth; occipital carina strong but incomplete ventromedially; mandible narrowest submedially, tapering apically toward ventroapical tooth.

Mesosoma. Pronotum nearly vertical anteriorly, punctures deep, contiguous close, shoulders well-developed, not produced laterally; foretibia without spines except apically; basitarsus with row of three spines on outer surface; tibial spur 0.9 × length of basitarsus; hindcoxal longitudinal dorsal carina elevated, angulate basally; mesonotal and mesopleural punctures deep, 0.5-1.0 puncture diameter apart; scutellum elevated above level of propodeum, with polished impunctate area medially; postscutellum densely punctate; propodeum nearly impunctate, dorsal surface with shallow medial, longitudinal groove, dorsal surface separated from posterior surface by transverse ridge, ridge interrupted medially, lateral surface with punctures 1-2 puncture diameters apart; forewing with 3 submarginal cells; costa continuing along wing margin beyond stigma.

Metasoma. Tergum I without transverse carina; sternum II without raised area basally.

Genitalia ( Figs 43 , 51 ). Gonostylar apex arcuately excavate in lateral view; aedeagus not decurved before apex.

Color. Head and body concolorous castaneous to black; vestiture brown.

Female unknown.

Distribution

( Fig. 37 ).

Etymology.

The species is named after the collector, John Chemsak.

Figures

Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance.

Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3.

Figures 37 - 40. Distribution maps Colocistis species.

Figures 41 - 48. Ventral view of male genital capsule.

Figures 49 - 56. Lateral view of male genital capsule.