Clidicus kalis Jałoszyński, Cheng & Yin, 2019

Jałoszyński, Paweł, Cheng, Zhi-Fei, Brajczewska, Nastazja & Yin, Zi-Wei, 2019, Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4658 (3), pp. 571-584 : 576-578

Clidicus kalis Jałoszyński, Cheng & Yin sp. n.

( Figs 14–20 )

Material studied. HOLOTYPE: ♂: PHILIPPINES, with two labels: “ PHILIPPINES / 6°55’N 125°16’E / Min- danao, prov. Davao der Sul / Kapatagan / IV 2018 / leg. Ismael Lumawig (local coll.)” [white, printed], “ Clidicus / kalis Jałoszyński et al. / HOLOTYPUS ” [red, printed] ( USMB) . PAPATYPES (19 ♂♂, 29 ♀♀): PHILIPPINES: 1 ♀, same data as for holotype ; 1 ♂, 2 ♀♀, “ PHILIPPINES / 07°39’N 124°42’E / Mindanao, prov. Lanao der Sul / Wao / III 2018 / leg. Ismael Lumawig (local coll.)” [white, printed] ; 1 ♂, same data except for XI 2018 ; 1 ♀, same data except for IX 2018 ; 1 ♂, same data, except for II 2019 (paratypes in USMB and cPJ) ; 1 ♂, same data as for holotype, except for XI 2017 ( SNUC) ; 1 ♂, 4 ♀♀, “Philippines / Mindanao / prov. Lanao del Sur / Wao / XII 2016 / leg. Ismael Lumawig (local coll.)” [white, printed] ( SNUC) ; 2 ♂♂, same data except for I 2017 ( SNUC) ; 2 ♀♀, same data except for II 2017 ( SNUC) ; 1 ♂, 3 ♀♀, same data except for IV 2017 ( SNUC) ; 1 ♂, 1 ♀, same data except for VII 2017 ( SNUC) ; 1 ♀, same data except for VIII 2017 ( SNUC) ; 2 ♂♂, 3 ♀♀, same data except for X 2017 ( SNUC) ; 1 ♂, 10 ♀♀, same data except for VI 2018 ( SNUC) ; 1 ♂, same data except for VIII 2018 ( SNUC) ; 1 ♀, same data except for XI 2018 ( SNUC) ; 2 ♂♂, “ Philippines / Mindanao / prov. Bukidnon / Kabanglasan / XII 2014 / leg. Ismael Lumawig (local coll.)” [white, printed] ( SNUC) ; 4 ♂♂, “Philippines / N. Luzon / Nueva Vizcaya / Kayapa / I 2017 / leg. Ismael Lumawig (local coll.)” [white, printed] ( SNUC).

Diagnosis. Pronotum with unevenly distributed, inconspicuously small, shallow and sparse punctures (distances between most punctures larger than their diameters), at middle of disc with impunctate or nearly impunctate small irregular oval area or line, cuticle on pronotum between punctures glossy, pronotal disc with a narrow, shallow and diffuse, barely noticeable median longitudinal impression in posterior 1/6–1/4; aedeagus with asymmetrical parameres, both in ventral view distinctly undulated, the right one with its apex indistinctly curved mesad.

Description. Body of male ( Fig. 14 ) elongated and slender, head flattened, prothorax and elytra convex; legs and antennae long and slender, BL 6.41–6.88 mm (mean 6.63 mm); cuticle glossy on head and pronotum, slightly less so on elytral disc, pigmentation uniformly reddish brown, setae indistinctly lighter than cuticle.

Head ( Fig. 15 ) broadest near middle of tempora, HL 1.05–1.18 mm (mean 1.12 mm), HW 1.43–1.50 mm (mean 1.47 mm); tempora in dorsal view approximately 3 times as long as eyes, in anterior third straight and distinctly di- vergent posterad, posteriorly evenly, strongly rounded; vertex with a deep subtriangular posteromedian impression and a pair of shallow pits marking the sites where dorsal tentorial arms are fused with the head wall, sides of vertex convex; frons between antennal insertions nearly flat, vertex and posterior portion of frons with narrow median longitudinal groove, which is more pronounced posteriorly; supraantennal tubercles small but distinctly elevated; compound eyes small, nearly round, in dorsal view indistinctly projecting laterally from the head silhouette, finely faceted. Punctures on sides of vertex and frons distinct but shallow and with diffuse margins, except for groups of several densely distributed punctures near mesal margin of each eye and posterolateral margin of each supraantennal tubercle; posteromedian impression on vertex, median line on frons and vertex, and supraantennal tubercles indis- tinctly punctate or impunctate. Setae long, sparse, suberect to erect. Antennae ( Fig. 16 ) slender, all antennomeres elongate (antennomere XI indistinctly), scape slightly longer than head, AnL 3.88–4.29 mm (mean 4.08 mm).

Labrum with straight anterior margin bearing an indistinct, barely noticeable median notch.

Pronotum with subpentagonal disc and short posterior ‘collar’, broadest near anterior third; PL 1.50–1.74 mm (mean 1.61 mm), PW 1.45–1.60 mm (mean 1.52 mm); ‘collar’ demarcated by a narrow transverse groove with 6–8 diffuse pits; additionally, two or three larger pits are present on each side of pronotum in its constricted portion. Punctures on disc distinct but small, shallow and diffuse, unevenly distributed, with variable spaces between punctures, but most punctures are separated by spaces wider than their diameters; elongate median area or line nearly impunctate or at least with punctures sparser than on sides; setae similar to those on head but indistinctly thicker.

Elytra approximately shield-shaped, broadest at or indistinctly behind middle, EL 3.67–4.05 mm (mean 3.90 mm), EW 2.15–2.40 mm (mean 2.28 mm), EI 1.61–1.81; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits slightly less glossy than on head and pronotum, between rows with very fine and sparse setiferous punctures. Apex of each elytron obliquely truncated, with moderately distinct, obtuse-angled lateral corner and distinct adsutural corner, which is strongly projecting posterad, sharp-angled and pointed. Setae on elytra distinctly longer and thicker than those on pronotum, suberect to erect.

Hind wings absent.

Legs long and slender, unmodified.

Aedeagus ( Figs 17–18 ) strongly elongate but moderately slender, AeL 1.45–1.55 mm (mean 1.52 mm); para- meres of unequal length and shape, in ventral view both undulated and with similar, slightly curved mesad apices, parameres distinctly divergent distad; flagellum differentiated from ductus ejaculatorius by a rapid broadening in proximal third of median lobe. ‘Sperm pump’ ( Fig. 19 ) 0.94–1.08 mm (mean 0.99 mm) in length, with two large ‘funnels’, distal funnel much shorter and slightly brooder than proximal one.

Female. Externally very similar to male, on average smaller and with shorter antennae; BL 6.33–6.92 mm (mean 6.59 mm); HL 1.10–1.18 mm (mean 1.15 mm), HW 1.53–1.70 mm (mean 1.61 mm), AnL 3.58–4.05 mm (mean 3.81 mm); PL 1.51–1.74 mm (mean 1.63 mm), PW 1.45–1.60 mm (mean 1.51 mm); EL 3.63–4.10 mm (mean 3.81 mm), EW 2.23–2.45 mm (mean 2.36 mm), EI 1.52–1.70.

Spermatheca ( Fig. 20 ) with ovoid capsular part, elongate, broadened and slightly irregular in shape projection connected to ductus spermathecae, width of capsular part 0.125 mm; accessory gland with connecting duct con- spicuously long; ductus spermathecae moderately strongly entangled.

Distribution. South-central and north-central Mindanao, northern Luzon.

Etymology. The name refers to undulated parameres which resemble the kalis, a traditional Philippine sword (the name is a noun in apposition).

Remarks. This species is most similar to C. interfector ; see remarks for that species. Also in C. kalis the length of parameres is variable between local populations.

Figures

FIGURES 14–20. Clidicus kalis Jałoszyński et al. sp. n. Habitus of male (14); head of male in dorsal view (15), right antenna of male in dorsal view (16); aedeagus in ventral (17) and lateral (18) views; ‘sperm pump’ (19); spermatheca (20).