Cladiopsocus sixtoi, Obando & García Aldrete & Carrejo, 2018

Obando, Ranulfo González, García Aldrete, Alfonso N. & Carrejo, Nancy, 2018, New species and records of Cladiopsocus Roesler (Psocodea: ‘ Psocoptera’: Psocomorpha: Cladiopsocidae) from Colombia, Zootaxa 4378 (2), pp. 191-223 : 209-213

Cladiopsocus sixtoi n. sp. Male

( Figs 76–82 )

Diagnosis. Belonging in species group “1”, subgroup 2 in the classification of Eertmoed (1986). The wings, paraprocts and phallosome relate it to C. eertmoedi González, García Aldrete & Carrejo , but unlike it the forewings are strongly pigmented, the inner arm of the paraproctal prong is almost as long as the outer arm ( Fig. 82 ), and the epiproct has a median projection rounded posteriorly ( Fig. 81 ).

Color. Head ( Fig. 78 ) brown, vertex, occiput and front brown, labrum, labium, mandibles and maxillae pale brown. Compound eyes dark brown, ocelli hyaline in a dark brown area. Antennae and maxillary palps pale brown, flagellomeres cream distally. Pronotum pale brown, mesonotum dark brown, metanotum light brown, with mesal line ochre, extended to metascutellum; thoracic pleura and legs light brown, coxae, tibia and tarsi darker. Forewing membrane pigmented, with brown-smoked areas and a brown band from R4+5 to CuA2 ( Fig. 76 ); veins brown, with pale brown spots along the veins; wing margin with brown spots. Hindwings hyaline to pale brown, as illustrated ( Fig. 77 ). Abdomen pale brown, with incomplete abdominal segmental rings pale brown, clunium, hypandrium and phallosome brown; epiproct cream; paraprocts pale brown, paraproctal prong brown, darker distally.

Morphology. As in diagnosis plus the following: head vertex emarginate, almost at the same level of the upper margin of the compound eyes ( Fig. 78 ); these large, with interommatidial setae. D is 0.57 the length of the head in front view. Forewings ( Fig. 76 ); membrane pterostigma with setae, with spur veins in areola postica. Hypandrium triangular, with medium sized setae and macrosetae. Phallosome of two lobes anteriorly, distinctly separated, distally wider, external parameres slender, membranous, bearing pores; endophallic lobes broadly triangular, separate along the midline; arms of internal parameres slender, bow shaped ( Fig. 80 ). Epiproct semioval, broad, with widened medial projection, rounded posteriorly ( Fig. 81 ), with three macrosetae and two setae mesally, posterior setal field as illustrated. Paraprocts with short mesal tubercle, with abundant medium setae and macrosetae as illustrated ( Fig. 82 ); mesal prong stout, sclerotized, tapered, of two arms, the inner arm almost as long as the outer one; sensory fields with 18 trichobothria on basal rosettes ( Fig. 82 ).

Measurements. FW: 3450, HW: 2600, F: 800, T: 1425, t1: 550, t2: 60, t3: 77, ctt1: 28, f1: 650, f2: 510, f3: 412, f4: 345, f5: 230, f6: 175, f7: 162, Mx4: 230, IO: 220, D: 453, d: 320, IO/d: 0.69, PO: 0.71.

Specimens studied. Holotype male. COLOMBIA. Caquetá . San Vicente del Caguán, Laureles, Resguardo Indígena Altamira, 2°27’50.14’’N : 74°55’02.06’’W, 917 m. 26–27.III.2017. J. Panche. Led light trap in forest canopy. MUSENUV slide code 2896. Paratypes: 3 males, same data as the holotype .

Etymology. This species is dedicated to Sixto Muñoz, the last of the Tinigua, an indigenous ethnic group that lived in an extensive area of the Colombian department of Caquetá.

Figures

FIGURES 76–82. Cladiopsocus sixtoi n. sp. Male. 76. Forewing. 77. Hindwing.78. Front view of head. 79. Hypandrium.80. Phallosome. 81. Epiproct.82. Paraproct.Scales in mm.