Cimrmaniela smoljaki, Frýda, Jiří, Ferrová, Lenka & Frýdová, Barbora, 2013

Frýda, Jiří, Ferrová, Lenka & Frýdová, Barbora, 2013, Review of palaeozygopleurid gastropods (Palaeozygopleuridae, Gastropoda) from Devonian strata of the Perunica microplate (Bohemia), with a re-evaluation of their stratigraphic distribution, notes on their ontogeny, and descriptions of new taxa, Zootaxa 3669 (4), pp. 469-489 : 474-476

Cimrmaniela smoljaki sp. nov.

Holotype. UUG JF 907 ( Figure 2 G–I, M).

Paratypes. UUG JF 908–913 ( Figure 2 J–L and 5 additional shells). Type locality. Císařský lom quarry (49°55'00.99"N, 14°3'36.420"E), S of the village of Konĕprusy. Type horizon. Middle part of the Konĕprusy Limestone, Praha Formation. Etymology: smoljaki , in honour of Ladislav Smoljak (1931–2010), one of the preeminent biographers of Jára

Cimrman.

Diagnosis: Species of Cimrmaniela ornamented by opisthocline costae, having regularly arched whorl side, narrow pleural angle.

Description: High-spired, narrow, multiwhorled, dextrally coiled shells, with up to 13 whorls; shell sides straight with pleural angle of 7 degrees; apical angle of first three whorls about 10 degrees; lateral whorl sides very convex, symmetrically arched; sutures relatively deep; sutural slope angle about 22 degrees; base of whorls rounded, anomphalous; external surface of whorls ornamented by regularly spaced costae; beginning at lower suture, costae run in opisthocline direction (also parallel with shell axis) across lateral whorl side, only just below upper suture gradually curving in an orthocline, then in prosocline direction; distance of costae about double their width ( Fig. 2 J–M); diameter of first half of shell whorl about 0.26 mm; first costae preserved from end of second whorl; preservation of shell not allowing for determination of exact position of protoconch/teleoconch boundary.

Comparison: Shells of C. smoljaki sp. nov. are ornamented by opisthocline costae, in contrast to C. sveraki sp. nov. (the only other species of Cimrmaniela ) shells, which bear orthocline costae. In addition, the sides of whorls in C. smoljaki are regularly arched, but the whorl side in C. sveraki is distinctly flattened below the upper suture, and strongly arched in a mid-whorl position. The apical angle of first three whorls is about 10 degrees in C. smoljaki , and is much wider in C. sveraki - about 22 degrees ( Fig. 3 ).

Stratigraphic and geographic distributions. Cimrmaniela smoljaki is only known from its type locality and horizon.

Cimrmaniela sveraki sp. nov.

Holotype. UUG JF 914 ( Figure 2 A–D).

Paratypes. UUG JF 915–921 ( Figure 2 E–F, and 6 additional shells).

Type locality. Císařský lom quarry, S of the village of Konĕprusy (49°55'00.99"N, 14°3'36.420"E).

Type horizon. Middle part of the Konĕprusy Limestone, Praha Formation.

Etymology: sveraki , in honour of Zdenĕk Svĕrák, one of the preeminent biographers of Jára Cimrman.

Diagnosis: Species of Cimrmaniela ornamented by orthocline costae; whorl side is distinctly flattened below the upper suture and strongly arched in mid-whorl position; diameter of embryonic shell about 0.17 mm, and diameter of the first half of whorl about 0.30 mm.

Description: High-spired, multiwhorled, dextrally coiled shell, with up to nine whorls present; shell sides slightly convex with pleural angle 9 degrees; initial shell having more convex sides than adult shell; apical angle of first three whorls about 22 degrees; lateral whorl sides distinctly flattened below upper suture, strongly arched in mid-whorl position; sutures relatively deep; sutural slope angle in juvenile shell about 13 degrees, about 17 degrees on adult whorls; base of whorls rounded, anomphalous; aperture circular; external surface of whorls ornamented by regularly spaced, orthocline costae; distance of costae about double their width ( Fig. 2 ); diameter of first shell whorl about 0.30 mm; diameter of large embryonic chamber about 0.17 mm; first costae preserved on second whorl; preservation of shell not allowing for determination of exact position of protoconch/teleoconch boundary.

Comparison: Differences of both of the known species of Cimrmaniela are described in the section on C. smoljaki .

Remarks. The protruding and large embryonic chamber in C. sveraki ( Fig. 2 E–F) fits well in the morphological range of the protoconchs of other palaeozygopleurid genera.

Stratigraphic and geographic distributions. Cimrmaniela sveraki is only known from its type locality and horizon.

Figures

FIGURE 2. Cimrmaniela sveraki sp. et gen. nov. (A – F) and Cimrmaniela smoljaki sp. et gen. nov. (G – M) from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A – D. Holotype (UUG JF 914), E – F. Paratype A (UUG JF 915). G – I and M. Holotype (UUG JF 907), J – L. Paratype (UUG JF 908). Shell height: A – D— 3.2 mm, E, F— 2.1 mm, H, I— 4.2 mm, J – L— 2.1 mm, figure height of shell detail: G— 1.9 mm, M— 1.8 mm.

FIGURE 3. Palaeozygopleura lukesi sp. nov. from middle Early Devonian (Pragian) age strata of the Konĕprusy Limestone (Praha Formation) in Barrandian, Czech Republic. A – C, H, and I. Holotype (UUG JF 922), D. Paratype A (UUG JF 923), E, F. Paratype B (UUG JF 924), G. Paratype C (UUG JF 925), J, M. Paratype D (UUG JF 926), K. Paratype E (UUG JF 927), L, N. Paratype F (UUG JF 928). All samples are from the type locality. A— shell width 1.5 mm, shell height: B, C— 4.6 mm, D— 3.9 mm, E, F— 3.8 mm, K— 3.6 mm, L— 5 mm, figure height of shell detail: G— 1.1 mm, H— 1.5 mm, I— 1.6 mm, J— 5.6 mm, M— 2.8 mm, N— 2.1 mm.