Chlamydastis ungulifera ( Meyrick, 1929 ), 1955

Phillips-Rodríguez, Eugenie, Brown, John W., Hallwachs, Winnie & Janzen, Daniel H., 2021, Chlamydastis Meyrick of Costa Rica: barcodes, biology, and descriptions of 36 new species (Lepidoptera: Depressariidae), Insecta Mundi 2021 (868), pp. 1-96 : 46

Chlamydastis ungulifera ( Meyrick, 1929)

Figures 49 , 88 , 122

Ptilogenes ungulifera Meyrick 1929: 517 .

Chlamydastis ungulifera: Clarke 1955: 207 .

Holotype. Male , Colombia, Gorgona Island, 200′, at light, 20.xi.1924. St. George expedition, C.L. Collenette, JFGC Slide No. 5614 ( NHMUK).

Specimens examined (51♂, 7♀). See Appendix 1.

Diagnosis. Superficially, C. ungulifera can be distinguished from Costa Rican congeners by the combination of two forewing features: a dark, crescent-shaped mark at the termen and a more rounded apex. The most conspicuous feature of the male genitalia that distinguishes it from congeners is the short, apically truncate uncus with a rounded basal portion.

Redescription. MALE ( Fig. 49 ). Head. Frons mostly whitish with two dark spots between antennal base and base of haustellum, vertex beige with pale brown scales; labial palpus beige; antenna with sensory setae ca. 1.5 times width of flagellomere. Thorax. Tegula and dorsum beige with brown line at base of tegula and collar, four distinct darker spots in center. FW length 7.0– 7.7 mm; FW ground color beige; termen with ovate blotch of pale brown scales bordered basally by a dark brown crescent; a small diffuse semicircular patch at costa ca. 0.15 from base to apex; a larger dark brown dash from costa just before apex. HW pale olive brown. Abdomen. Pale brown externally. Genitalia ( Fig. 88 ) with uncus short, truncate distally, with rounded base; gnathos with long, triangular plate at distal junction of arms; tegumen slightly sclerotized anterior to base of uncus, with short portion of posterior border strongly sclerotized; lateral processes of juxta bladelike, sclerotized; valva with specialized setae from a rounded subapical patch; phallus simple with a crescent-shaped, dentate cornutus.

FEMALE. Head and Thorax. Essentially as described for male, except sensory setae of antenna short, sparse; FW length 7.1–7.3 mm. Abdomen. Genitalia ( Fig. 122 ) with outer margins of papillae anales slightly diverging posteriorly; sterigma a simple band with sclerotized area around ostium; ductus bursae short, wide; corpus bursae ovate with small, weakly dentate signum.

DNA barcodes. Sequence data are unavailable for this species.

Distribution. Chlamydastis ungulifera was described from Gorgona Island, Colombia; we recorded it in Costa Rica from 0 to 200 m elevation in the central and south Pacific coasts.

Biology. The immatures and food plant are unknown.

Figures

Figures 42–50. Adults of Chlamydastis from Costa Rica. 42) C. irenecanasae, USNMENT01480480. 43) C. dondavisi, USNMENT01480468. 44) C. manuelzumbadoi, female, paratype, INBIOCRI001956652. 45) C. noramartinae, male, paratype, INBIOCRI000598917. 46) C. vitorbeckeri, male, holotype, 11-SRNP-43642. 47) C. ronaldzunigai, holotype, 11-SRNP-20864. 48) C. munifigueresae, 12-SRNP-41409. 49) C. ungulifera, male, INBI- OCRI002540131. 50) C. willsflowersi, male, paratype, INB0003901728.

Figures 83–89. Male genitalia (phallus detached) of Chlamydastis from Costa Rica – Cystiodes, Ronaldzunigai, Unassigned Species Groups. 83) C. manuelzumbadoi, paratype, 08-SRNP-70120. 84) C. noramartinae, holotype, 11-SRNP-104804. 85) C. vitorbeckeri, paratype, 09-SRNP-73023. 86) C. ronaldzunigai, paratype, IN- BIOCRI000306854. 87) C. munifigueresae, paratype, 12-SRNP-41409. 88) C. ungulifera, INB001969967. 89) C. willsflowersi, paratype, 09-SRNP-106691.

Figures 118–123. Female genitalia of Chlamydastis. 118) C. manuelzumbadoi, paratype, 08-SRNP-70122. 119) C. vitorbeckeri, paratype, 09-SRNP-44277. 120) C. ronaldzunigai, paratype, INBIOCRI000306853 (detail of signum, INBIOCRI0001747283). 121) C. munifigueresae, paratype, 10-SRNP-42832. 122) C. ungulifera, INBI- OCRI000445676. 123) C. willsflowersi, paratype, INBIOCRI003901728.