Chimarra kokodana Kimmins, 1962

Cartwright, David, 2020, A review of the New Guinea species of Chimarra Stephens (Trichoptera: Philopotamidae), Memoirs of Museum Victoria 79, pp. 1-49 : 35

Chimarra kokodana Kimmins, 1962

Figures 106, 107

Chimarra kokodana Kimmins, 1962: 119 ; figs 23, 24.— Neboiss, 1986a: 109.

Type material not seen. Holotype. Male , PNG, Kokoda, 1200 ft (about 366 m), August 1933, L.E. Cheeseman ( BMNH).

Material examined. PNG, Central Province. 1 male (dried, pinned specimen CT-356 partly figured), Mamai Plantation, east of Port Glasgow , 150 m, about 10° 16' S, 149° 30' E, lt tr, 29 January 1965, R. Straatman ( BPBM); 14 males (CT-345), Iomari Creek , Bereima – Port Moresby Rd, about 9° 25' S, 147° 15' E, UV light, 23 May 1986, A. Wells and W. Ismay ( NMV) .

Diagnosis. The males of C. kokodana are similar to C. bifida and C. bicuspidis in the bifid apices on the inferior appendages, but can be separated from the latter two and all other New Guinea species, by the combination of the inferior appendages, which are relatively straight and angled at between about 30° and 45° to horizontal in lateral view, with bifid apices and the rounded, keel-like ventral process on segment IX.

Description. (Revised after Kimmins, 1962). General body colour and wings fawn (personal observation) to ochraceous ( Kimmins 1962: p. 119). Wings ( Kimmins, 1962: fig. 23), similar to those of C. ukarumpana (fig. 7). Length of forewing: male 3.8–4.8 mm. Forewing with forks 1, 2, 3 and 5 present, Rs moderately sinuous or curved, strongly thickened, basal to discoidal cell ( Kimmins, 1962: fig. 23); hind wing with forks 1, 2, 3 and 5 present.

Male. Segment IX anterior margin in lateral view, anteroventrally rounded (fig. 106, Kimmins, 1962: fig. 24A), ventral process short, basal to distal margin of segment IX, in lateral view, keel-like with rounded distal margin, length about half basal width (fig. 106), preanal appendages, ovate (figs 106, 107). Segment X lateral lobes relatively long, aligned laterad to phallus, sensilla not discerned (figs 106, 107), in lateral view, appears tapered in distal third (fig. 106), in dorsal view slender, apices appear slightly dilated or bulbous (fig. 107; Kimmins, 1962: fig. 24B). Phallus with two slender spines embedded subapically. Inferior appendages slightly broader in basal third, apices directed posteromesally, bifid (figs 106, 107), in lateral view, angled at between about 30° ( Kimmins, 1962: fig. 24A) and 45° to horizontal, length about 3.4 to 3.7 times width, dorsal and ventral margins straight in distal two thirds (fig. 106), in dorsal view mesal and lateral margins curved (fig. 107; Kimmins, 1962: fig. 24B).

Female. Unknown (a female was referred to this species with some doubt [ Kimmins, 1962: p.121]).

Remarks. Chimarra kokodana is known from 17 male (and one female?) specimens from four localities in south-east PNG and New Britain (referred to as C. kokoda by Oláh and Mey, 2013, p. 413). New figures have been drawn to allow direct comparisons and to accompany the description that is revised in light of new interpretations of Chimarra genitalic structures from Kimmins’ (1962) original description.

Figures

Figures 103–110. Chimarra spp.; 103–105, Chimarra erecta sp. nov., male, holotype, genitalia; 103, lateral; 104, ventral; 105, dorsal. 106–107, Chimarra kokodana Kimmins, male, genitalia; 106, lateral; 107, dorsal. 108–110, Chimarra espelandae sp. nov., male, holotype, genitalia; 108, lateral; 109, ventral; 110, dorsal.