Ceratostigma lisae ( Thornton, 1984 ) Jie & Liang & Liu, 2022

Jie, Lulan, Liang, Feiyang & Liu, Xingyue, 2022, Revision of the bark louse genus Ceratostigma Li, 2002 (Psocodea: Psocidae: Psocinae) with description of a new species from China, Zootaxa 5168 (2), pp. 196-206 : 199

Ceratostigma lisae ( Thornton, 1984) comb. n.

Metylophorus lisae Thornton, 1984: 140 . Type locality: Indonesia (Bali).

Diagnosis. Male. Forewing radial fork with branches diverging at an angle of about 30°. Epiproct rounded at apex; hypandrium with a narrowed ridge medially. Female. Forewing radial fork with branches diverging at an angle of less than 90°; cells r3 and r5 almost hyaline; basal area with irregular hyaline areas. Pigmented area of subgenital plate roughly T-shaped, nearly pointed at apex; dorsal valve anterior one-third with a small slightly sclerotized area.

Distribution. Indonesia (Bali, Flores, Java).

Remarks. This species was originally placed in Metylophorus Pearman, 1932 (Metylophorini Roesler, 1943) because of its similar color of wings and venations with most species of Metylophorus . Metylophorus can be diagnosed by having the forewing veins entirely brown; hypandrium broad, divided into three lobes, median lobe strongly sclerotized, usually with strongly sclerotized projections; subgenital plate with a long egg guide; dorsal valve of gonapophyses with a round apex. The type species of Metylophorus , M. nebulosus (Stephens, 1836) , can be separated from C. lisae by male hypandrium asymmetrical, divided into three lobes, median lobe with two left processes well developed and two small right processes close to the base; female egg guide of subgenital plate elongate ( Endang et al. 2002; Li 2002; Kido & Yoshizawa 2019; Pearman 1932; Thornton 1984). C. lisae can be distinguished from M. nebulosus by having the forewing with veins on the basal two-thirds yellowish or grayish white; male hypandrium a simple plate, medially with a ridge; female subgenital plate with a short egg guide; dorsal valve of gonapophyses broad, pointed at tip. Thus, it seems reasonable to transfer this species to Ceratostigma , and the new combination proposed above. Prior to this study, all the species of Ceratostigma were only known from China, while the new combination represents the first record of this genus from Indonesia. Ceratostigma lisae can be distinguished from congeners by having the male forewing radial fork with branches diverging at an angle of about 30°, epiproct with a rounded apex, hypandrium medially with a narrowed ridge, female forewing radial fork with branches diverging at an angle less than 90° and cell r3 and r5 almost hyaline.