Cephennium (Macroderus) segurae, Assing, 2021

Assing, Volker, 2021, On the Cephennium fauna of the Iberian Peninsula and the Atlantic Islands (Coleoptera, Staphylinidae, Scydmaeninae), Linzer biologische Beiträge 52 (2), pp. 891-931 : 927

Cephennium (Macroderus) segurae nov.sp. ( Figs 17 , 24 , 67-69 , Map 3 )

Type material: Holotype ♂: "E – Castilla-La Mancha, Sierra de Segura, Calar de Sima , 1430 m, [3], 38°20'36N 02°28'31W, 15.III.2008, V. Assing / Holotypus ♂ Cephennium segurae sp. n. det. V. Assing 2020" ( cAss) . Paratypes: 2♂♂, 3♀♀: same data as holotype ( cAss) .

Etymology: The specific epithet is the genitive of Segura, the mountain range where the species is probably endemic.

Description: Body length 1.3-1.4 mm. Habitus as in Fig. 17 . Colouration reddish. Pubescence of pronotum and elytra moderately long and depressed. Eyes composed of approximately seven ommatidia. Pronotum approximately 1.35 times as broad as long. Elytra with anterior sulci approximately as long as the combined length of the basal four antennomeres. Other external characters as in C. alacantae .

♂: metaventrite shallowly impressed; aedeagus ( Figs 67-69 ) 0.32-0.33 mm long; apex of median lobe broadly obtuse in ventral view; internal structures large and distinctive; parameres apically nearly reaching apex of median lobe.

Comparative notes: Cephennium segurae differs from the geographically close C. cujonicum by slightly larger size, slightly darker colouration, a slightly more transverse pronotum, a shallowly impressed male metaventrite, and by the shape and internal structures of the aedeagus.

Distribution and natural history: The type locality is situated in the western Sierra de Segura range ( Map 3 ). The specimens were sifted from grass, moss, deep litter, and ferns between rocks in a deep limestone canyon with hazelnut at an altitude of 1430 m ( Fig. 24 ).

Figures

Map 3: Distributions of Cephennium asturicum (black circles; based on Besuchet (unpubl.) and examined material) and of species of the subgenus Macroderus in the Iberian Peninsula: C. alacantae (black triangles); C. cujonicum (white diamond); C. segurae (black diamond); C. verrens (white circles); C. cazorlae (black square); C. nevadae (black star); C. malagae (white triangles).

Figs 17-20: Male habitus of species of the subgenus Macroderus in dorsal view: (17) Cephennium segurae; (18) C. nevadae; (19) C. cazorlae; (20) C. malagae. Scale bar: 1.0 mm.

Figs 23-24: Type localities of Cephennium cujonicum (23; above) and C. segurae (24; below).

Figs 65-71: Cephennium cujonicum (65), C. verrens (66), C. segurae (67-69), and C. malagae (70- 71): (65-71) aedeagus in lateral and in ventral view. Scale bars: 65-71: 0.1 mm.