Cancilla wagreichi, Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 62

Cancilla wagreichi nov. sp.

Figs 15A 1 –A 2

[ Mitra ] tenuistria Duj. —Hoernes 1880: 125 [non Dujardin, 1837].

Mitra tenuistria Duj. — Hoernes & Auinger 1880: 80, pl. 9, figs 8a–b [non Dujardin, 1837].

Type material. Holotype: NHMW 2020 /0130/0001, SL: 29.3 mm, MD: 8.7 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger 1880, pl. 9, figs 8a–b, Figs 15A 1 –A 2 .

Type locality. Lăpugiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, early/middle Badenian (Langhian).

Etymology. In honour of Michael Wagreich (University Vienna), in respect for his contributions on Paratethyan stratigraphy.

Diagnosis. Cancilla species of medium size, slender fusiform profile, with moderately high spire, weakly convex whorls, elongate last whorl with shallow basal concavity, twisted siphonal canal, and delicate spiral sculpture of numerous flattish spiral cords.

Description. Shell medium sized, slender fusiform. Protoconch and early teleoconch unknown. Spire whorls weakly convex, with periphery in abapical third; suture narrowly incised. Last whorl elongate, evenly convex, slowly contracting into moderately long base. Sculpture consisting of six to seven broad spiral cords, with flat tops on early spire whorls. Number of cords increases to about 22 on penultimate whorl by intercalation of secondary spiral grooves. Spiral sculpture persists on last whorl, being most prominent below adapical suture and on base, but slightly blurred along periphery. Aperture elongate, columellar callus indistinct. Columella with four delicate columellar folds; outer lip thin. Fasciole weakly swollen. Siphonal canal short, moderately wide, weakly twisted.

Shell measurements and ratios. SL = 29.3 mm, MD: 8.7 mm; AA = 33°, SL/MD: 3.4, AL/AW: 4.7.

Discussion. The specimen from Lăpugiu de Sus was identified by Hoernes & Auinger (1880) as Mitra tenuistria Dujardin, 1837 , originally described from the middle Miocene of the Touraine ( France). The French species, however, is readily distinguished by its broader shell and the distinctly broader and more prominent spiral cords (see Dujardin 1837, pl. 20, fig. 26; Peyrot 1938, pl. 4, figs 32, 40; Glibert 1952b, pl. 12 fig. 3). We are aware of few species that are comparable to the Paratethyan species: Mitra bellatula Bellardi, 1887 , from the late Miocene of Stazzano ( Italy) has almost identical sculpture, but differs in its wider spire angle, broader last whorl and Mitrella -like shape (see Bellardi 1887a, pl. 4, fig. 2; Ferrero-Mortara et al. 1981, pl. 44, figs 7a–b). Mitra interposita Bellardi, 1887 (see Bellardi, 1887a, pl. 4, fig. 21; Ferrero-Mortara et al. 1981, pl. 49, figs 3a–b) is higher spired, has higher spire whorls, and coarser spiral sculpture. Mitra subuliformis Bellardi, 1887 , from the early Miocene of the Colli Torinesi ( Italy), is very similar in outline, but is smaller (SL = 20 mm), has a weaker fasciole and lacks the delicate spiral sculpture (see Bellardi 1887a, pl. 3, fig. 42; Ferrero-Mortara et al. 1981, pl. 46, figs 14a–b).

Palaeoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1880).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 15. A1–A2. Cancilla wagreichi nov. sp., holotype, NHMW 2020/0130/0001, Lăpugiu de Sus (Romania). B1–B2. Cancilla sismondai (Michelotti, 1847), NHMW 1865/0001/0171, Lăpugiu de Sus (Romania). C1–C2. Cancilla nanostriatula nov. sp., NHMW 2020/0136/0002, paratype, Bad Vöslau (Austria). D1–D2. Cancilla nanostriatula nov. sp., NHMW 1846/0037/0100, holotype, Baden (Austria). E1–E2. Cancilla nanostriatula nov. sp., NHMW 1863/0015/0622a, paratype, Niederleis (Austria). F1–F2. Cancilla nanostriatula nov. sp., paratype, Baden (Austria). G1–G2. Cancilla suballigata (Bellardi, 1887), NHMW 1847/0037/0044b, Vienna/Pötzleinsdorf. H1–H2. Cancilla suballigata (Bellardi, 1887), NHMW 2020/0137/0001, Bad Vöslau (Austria). I1–I2. Cancilla suballigata (Bellardi, 1887), NHMW 2020/0134/0001, Forchtenau (Austria).