Brachylinga bicolor Webb, 2006

Webb, Donald W. & Metz, Mark A., 2006, A Revision of the New World Genera Brachylinga Irwin and Lyneborg and Lysilinga Irwin and Lyneborg (Diptera: Therevidae: Therevinae) with the Description of a New Genus, Elcaribe Webb, Zootaxa 1288 (1288), pp. 1-241 : 52-55

Brachylinga bicolor Webb spec. nov.

( Figs. 6, 29, 52, 75, 98, 121, 144, 167, 188, 217)

Etymology

Bi (Latin) = two, twice; color (Latin) = color. This name is used as an adjective and refers to the dark and light coloration of the male parafacial.

Diagnosis

Brachylinga bicolor is taxonomically associated with B. divaricata in lacking setae on the katepisternum; in having the postocular, mesonotal, and coxal macrosetae dark reddish brown; in having the male parafacial black ventrally, silver dorsally; and in having the hypoproct quadrate and the posterior margin slightly emarginate. Brachylinga bicolor differs from B. divaricata in having the gonocoxite ( Fig. 29) oblong, tapered apically and lacking short, dark reddish brown macrosetae apicomedially and elongate, dark reddish brown macrosetae anteromedially. Females of B. bicolor , B. concava , B. divaricata , and B. ornata are indistinguishable based on external characters. Internally B. bicolor , B. concava and B. ornata have the spermathecal ducts directed anteriorly ( Figs. 188, 192, 199) while in B. divaricata the spermathecal ducts are directed laterally before bending at a right angle and continuing anteriorly ( Fig. 194).

Description of holotype male (MEI 007477)

Body length 7.0 mm.

Head. Length 0.94 mm. Ommatidia smaller ventrally and laterally. Frons silver, dark brown dorsally; setae black. Antenna/head length 0.69; scape and pedicel dark reddish brown, pruinescence gray, flagellum dark brown; scape length 0.21 mm, width 0.14 mm, length/width 1.5, scape/pedicel length 2.7, scape/flagellum width 0.74, setae pale brown, elongate; pedicel length 0.08 mm, length/width 0.6; flagellum length 0.36 mm, width 0.19 mm, length/width 1.9, flagellum/scape length 1.7 times. Parafacial pruinescence black, dorsal half silver. Maxillary palpus dark reddish brown; length 0.42 mm, length/width 7.0; setae pale yellow. Genal setae whitish yellow. Occipital setae whitish yellow, lanceolate dorsally. Postocular macrosetae dark reddish brown.

Thorax. Macrosetae dark reddish brown, 3 np, 2 sa, 1 pa, 2 dc, 2 sc. Mesonotum black; setae black, moderately long intermixed with white, lanceolate, subappressed setae. Pleuron pruinescence silver, dense; setae white on propleuron, anepisternum, laterotergite, metanepisternum, and scutellum, absent on katepisternum. Wing. Membrane pale gray; veins brown; length 5.3 mm, length/width 3.1; pterostigma brown, narrow; cell m 3 closed, petiolate. Halter pale yellow. Legs. Coxa setae white, present on posterior surface of midcoxa; macrosetae dark reddish brown. Femora dark reddish brown, pruinescence white; ventral setae black on fore­ and midfemora; av 0:0:4, pv 0:0:5. Tibiae dark yellow, apical fourth dark reddish brown. Tarsomere 1 dark yellow, apical half dark reddish brown; 2–5 dark reddish brown.

Abdomen. Dark reddish brown, pruinescence white, dense; dorsal setae white, appressed; lateral setae white, erect. Terminalia (MEI 121613) dark brown. Tergite 8 ( Fig. 6), narrow medially (0.04 mm); setae pale yellow, absent medially on posterior margin. Sternite 8 ( Fig. 29), sides rounded, posterior margin slightly emarginate. Epandrium ( Fig. 6), sides tapered apically, setae pale yellow. Cercus ( Fig. 6), ending basal to apex of hypoproct. Hypoproct ( Fig. 6), posterior margin emarginate, ending distal to apex of cercus. Gonocoxite ventral view ( Fig. 29), basal half quadrate, rounded laterally, posterior half attenuate, setae pale yellow (occasionally dark red), macrosetae dark brown apically; dorsal view ( Fig. 52) posteromedial lobe short, rounded, gonocoxal bridge absent; lateral view ( Fig. 75) broad basally, tapering apically as thick projection; ventral lobe ( Fig. 29) with ventral spines. Gonostylus ( Fig. 98), basodorsal lobe large, curved anteriorly; ventral margin straight, setae dark yellow, thick, extending apically in distinct tuft; fine, short spines absent laterally. Aedeagus with dorsal apodeme ( Fig. 121) narrow, rectangular, posterolateral margin rounded; ventral apodeme ( Fig. 144) tapered anteriorly, ending before anterior margin of dorsal apodeme; distiphallus dorsal view ( Fig. 121) tapered apically, lateral view ( Fig. 167) tapered apically in broad curved, apicolateral spines present; ejaculatory apodeme ( Fig. 121), anterior margin truncate.

Variation (n=10). Body length 5.2–6.3, 5.7 mm.

Thorax. Macrosetae 3 np, 2 sa, 1 pa, 2 dc, 2 sc. Wing. Cell m 3 widely open (MEI 106340). Legs. Anteroventral macrosetae 0:0:3–5, pv 0:0:2.

Female

Similar to male except for following.

Variation (n=10).

Body length 5.5–7.5, 6.2 mm.

Head. Frons pruinescence brownish gray, ventral fourth white to silver with large brown circle dorsolaterally; setae dark brown. Scape over 3 times length of pedicel, setae black, short, grading into macrosetae. Parafacial dark reddish brown, silver gray dorsally and laterally.

Thorax. Macrosetae 2–3 np, 2 sa, 1 pa, 2 dc, 2 sc. Mesonotal setae pale gold, lanceolate; scutellar setae pale gold. Legs. Anteroventral macrosetae 0:0:3–5, pv 0:0:4–8.

Abdomen. Coloration highly variable, generally dark reddish brown, tergites 2–6 with posterior margin dark yellow or yellowish orange, tergites 2­6 with posterolateral area of white priunescence. The abdomen may also be dark reddish brown with white pruinescence across posterior margin of tergite 2 and tergites 3–6 with posterolateral area of white priunescence; dorsal setae dark reddish brown on dark reddish brown areas and white appressed across posterior margin of tergites 1–3; lateral setae dark reddish brown. Terminalia (MEI 121593) reddish brown. Furca ( Fig. 188) oval; length 0.42 mm, width 24 mm; anterolateral projection absent. Common duct ( Fig. 188) shorter than furca. Spermathecal duct ( Fig. 188) directed anteriorly.

Distribution

Brachylinga bicolor is known from Costa Rica, Mexico, Nicaragua, and Venezuela ( Fig. 217).

Habitats and phenology

Brachylinga bicolor has been handnetted and collected in Malaise traps in rain forest or in pan traps. Specimens have been recorded during every month of the year except for July and October.

Specimens examined

Type specimen. The holotype male of Brachylinga bicolor Webb (MEI 007477) is labeled "COSTA RICA, Guanacaste Province, 10 km W San Miguel, 23 February 1991, DWWebb/MEIrwin, collected in Malaise trap" and is deposited in the INHS. PARATYPES. COSTA RICA. Guanacaste. Parque Nacional Santa Rosa, Liberia, Stor Santa Rosa [10.84, ­85.62], 300 m, –. I.1999, Sullivan, J., Perez, A. A., 11♀ INB 3067202, 3070483, 3070490, 3070683, 3070929–31, 3070934, 3070975–6, 3071042 ( INBC), Malaise trap, rain forest ; –. II.1999, 2♂ INB 3070920, 3073708, 13♀ 3073706–7, 3073710, 3073712–3, 3073835–6, 3073838–40, 3073842, 3073844–5 ( INBC), Malaise trap, rain forest ; –. III.1999, 1♂ INB 3071459, 36♀ 3068950–1, 3068954–60, 3068976, 3068981, 3071453–6, 3071458, 3071460, 3071843, 3071852–8, 3071860, 3071862, 3071864–5, 3071869–72, 3071874–5, 3071894 ( INBC), Malaise trap, rain forest ; –. IV.1999, 12♀ INB 3069210, 3069214, 3069218, 3069220, 3069223–5, 3069227–8, 3074275, 3074282, 3074284 ( INBC), Malaise trap, rain forest ; –. V.1999, 4♀ INB 3067196, 3068519, 3068525, 3068528 ( INBC); Parque Nacional Volcán Rincón de la Vieja, Estación de Las Pailas [10.78, ­85.35], 800 m , 10–18.IV.1994, Taylor, K., 1♀ INB 793001 ( INBC); 10 km SW San Miguel [10.336, ­85.014] , 25–27.III.1991, FDP, 4♂ MEI 047745, 047748–9, 047752 ( EMUS), Malaise trap; 10 km W San Miguel [10.4, ­85.04] , 21.II.1991, Webb, D. W., MEI, 5♂ MEI 007421, 007428, 007440, 007459, 007486 ( INHS), Malaise trap ; 22.II.1991, 10♂ MEI 007417, 007425, 007429–30, 007437, 007442, 007461, 007472, 007476, 011331 ( INHS) ; 23.II.1991 15♂ MEI 007412, 007419, 007422, 007427, 007439, 007448–9, 007452, 007456, 007468, 007478–9, 007493–4, 090008 ( INHS) ; 25.II.1991, FDP, 1♂ MEI 047530 ( EMUS), Malaise trap ; 28.II.1991, 2♂ MEI 047588, 047590

(EMUS); 19–20.III.1991, 3♂ MEI 047716, 047719, 047724 ( EMUS) ; 21–22.III.1991, 4♂ MEI 047722, 047725, 047730–1 ( EMUS) ; 23–24.III.1991, 6♂ MEI 047734, 047736, 047739–42 ( EMUS) ; 25–27.III.1991, 1♂ MEI 047757 ( EMUS) ; 1.IV.1991, 2♂ MEI 047770, 047772 ( EMUS) ; 8–12.IV.1991, 2♂ MEI 047698, 047700 ( EMUS) ; 16–23.II.1988, 1♂ MEI 072869 ( EMUS) ; 1–2.II.1989, 1♂ MEI 007855 ( EMUS) ; 2.III.1989, 1♂ MEI 036936 ( EMUS) ; 20–24.III.1989, 1♂ MEI 036916 ( EMUS) ; 21.III.1989, 1♂ MEI 072864 ( EMUS) ; 23–31.I.1990, 1♂ MEI 141753 ( EMUS) ; 1–5.II.1990, 1♂ MEI 036935 ( EMUS) ; 7–15.II.1990, 1♂ MEI 128863 ( INHS) ; 15–24.II.1990, 3♂ MEI 007860, 007870, 007874 ( EMUS) ; 16–22.II.1990, 1♂ MEI 141745 ( EMUS) ; 19–28.II.1990, 6♂ MEI 036905, 036918, 036920, 036922; 036930, 036933 ( EMUS) ; 23–28.II.1990, 11♂ MEI 007843–4, 007846–50, 007852, 007867, 036904, 036924 ( EMUS) ; 1–12.III.1990, 2♂ MEI 036910, 036921 ( EMUS) , 16♂ MEI 072916–21, 072923, 072932, 072935, 072937–8, 072940–1; 072943, 072945, 072947 (E MUS) ; 2.III.1990, 1♂ MEI 072876, 1♀ 072873 ( EMUS) ; 8–15.III.1990, 5♂ MEI 072862–3, 072880, 072898, 075283 ( EMUS) ; 9–14.III.1990, 1♂ MEI 128823 ( INHS) , 1♂ 141768 ( EMUS) ; 13–21.III.1990, 7♂ MEI 072874–5, 072886–7, 072906–7, 072909 ( EMUS) ; 15–22.III.1990, 1♂ MEI 036903 ( EMUS) ; 16–31.III.1990, 1♂ MEI 072955 ( EMUS) ; 22–31.III.1990, 7♂ MEI 072883, 072890, 072893–4, 072896, 072900–1 ( EMUS) ; 1–7.IV.1990, 2♂ MEI 007857, 128822 ( INHS) ; 24–31.VIII.1990, 1♂ MEI 036923 ( EMUS) ; 17–24.I.1991, 2♂ MEI 036913, 036919 ( EMUS) ; 1–7.II.1991, 2♂ MEI 036909, 036912 ( EMUS) , 3♂ 128820, 128839, 128875 ( INHS) ; 8–15.II.1991, 1♂ MEI 036907 ( EMUS) ; 16–23.II.1991, 1♂ MEI 036911 ( EMUS) ; 8–15.IV.1991, 1♂ MEI 036925 ( EMUS) ; 1–15.II.1992, 10♂ MEI 036906, 036908, 036914, 036917, 036927–9, 036931–2, 036934 ( EMUS) ; 15 km W San Miguel [10.4, ­85.085] , 23.II.1991, Webb , D. W., MEI, 1♂ MEI 007632 ( MEIC) ; 2 km SW del Cerro Cacao [10.93, ­85.47], 1000–1400 m , 8–17.II.1995, Zumbado, M. A., 1♀ INB 2323989 ( INBC) ; 3 km SE Rio Naranjo [9.912, ­84.951] , –. XII.1991, FDP, 1♂ MEI 141831 ( EMUS) ; 22–30.III.1992, 1♂ MEI 101267 ( EMUS) ; 15–30.IV.1992, 1♂ MEI 101194 ( EMUS) ; 24–31.XII.1992, 1♂ MEI 128843 ( INHS) ; 16–31.III.1993, 1♂ MEI 141839 ( EMUS) ; Cacao [10.93, ­85.47], 1100 m , 11–18.II.1995, Navarro, E., 1♀ INB 2184592 ( INBC) ; Los Alemendros [11.03, ­85.53], 300 m , 23–18.IV.1994, Lopez, E., 1♂ INB 1779775 ( INBC) ; S Cañas [10.43, ­85.1], 21–25.I.1989, FDP, 3♂ MEI 007826 ( BMNH) , 007853 (CSCA), 007854 (FSCA); 26–31.I.1989, 1♂ MEI 07856 ( USNM) ; 1–3.II.1989, 9♂ MEI 007830–1, 007833–7, 007840, 007863 ( EMUS) ; 9–14.II.1989, 3♂ MEI 007859, 007862, 007865 ( EMUS) ; 16–20.II.1989, 1♂ MEI 007858 ( EMUS) ; 1–5.III.1989, 1♂ MEI 007841 ( EMUS) ; 23–25.III.1989, 1♂ MEI 007839 ( EMUS) ; 4–6.V.1989, 1♂ MEI 007832 ( EMUS) ; 8–18.III.1988, 3♂ MEI 007824, 007828–29 ( EMUS) ; Santa Cruz , Quebrada Brazil, Estación B.N. Diria [10.17, ­85.6], 160 m , 12.III.2000, Cardenas , Y., 1♂ INB 3152607, 1♀ 3152606 ( INBC) , Malaise trap; Vista Del Mar [10.13, ­85.63], 972 m , 17–25.II.2002, Cardenas, Y., 1♂ INB 3441671 ( INBC) ; Tierras Morenas [10.57, ­85.03], 685 m , –. III.1994, Rodriguez , G., 2♂ INB 1683842, 1683848 ( INBC) ; Diria National Forest, Santa Cruz [10.13, ­85.63], 972 m , 16.III.2000, Cardenas, Y., 1♂ INB 3152743 ( INBC) ; 1.III.2001, Cardenas, Y., 1♀ INB 3131697 ( INBC) ; Palo Verde National Park, Bagaces , Cerro Guayacan [10.36, ­85.35], 195 m , 5–19.V.1999, Jimenez, I., 1♀ INB 3360680 ( INBC) , Malaise trap; Parque Nacional Rincon de la Vieja, Estación de Las Pailas [10.81, ­85.33], 800 m , 6–13.IV.1993, Sihezar, J., Rodriguez, G., 1♂ INB 331389 ( INBC) ; 800 m, 10–20.IV.1994, García, D., 1♀ INB 784421 ( INBC) ; Parque Nacional Barra Honda, Nacaome [10.217, ­85.35], 100 m , –. IV.1995, Reyes , M., 2♀ INB 2172192, 2172195 ( INBC) , yellow pan trap; Refugio Nacional de Fauna Silvestre Rafael Lucas Rodriguez , Palo Verde [10.35, ­85.35], 10 m , –. IV.1991, Chavarria , U., 1♂ INB 318929 ( INBC) ; Parque Nacional Santa Rosa , 300 m , –. III.1984, Janzen, D. H., Hallwachs, W., 1♂ MEI 048124 ( AMNH) ; –. IV.1984, MEI 048126 ( AMNH) ; 6.27. IX.1986, Janzen, D., Gauld, I., 1♂ MEI 138925 ( LACM) , Malaise trap; 21.II–14.III.1987, 14♂ MEI 138904, 138927, 138929, 110365, 122450–9 ( LACM) ; 14.III–4.IV.1987, 2♂ MEI 139013, 139020 ( LACM) , Malaise trap in treefall gap open to sun; Liberia, Stor Santa Rosa [10.84, ­ 85.62], 300 m, –. XI.1999, Janzen, 1♀ INB 3066755 ( INBC) , Malaise trap; Bagaces, Parque Nacional Palo Verde , 0.2 km NE Estación [10.37, ­85.38], 1–50 m , 6.III–6.IV.2000, Jimenez, I., 1♀ INB 3377263 ( INBC) , Malaise trap; Sector Palo Verde [10.37, ­85.38], 50 m , 5.XII.1999 – 5.I.2000, Jimenez, I., 1♀ INB 3369830 ( INBC) , Malaise trap; Playa Naranjo [10.78, ­85.67], 0 m , 11.VI.2001, Briceno, D., 1♀ INB 3332838 ( INBC) ; Canton de Bagaces, Parque Nacional Palo Verde , 0.2 km NE de la Estación, 1–50 m , 6.III–6.IV.2000, Jimenez, I., 1♂ INB 3377250 ( INBC) , Malaise trap; 0.2 km NE Estación [10.37, ­85.38], 0–50 m , 6.III–6.IV.2000, 1♀ INB 3377254 ( INBC) , Malaise trap; Cerro Guayacan, Sector Palo Verde [10.35, ­85.35], 212 m , 11.XI–8.XII.1999, 1♀ INB 3419745 ( INBC) , Malaise trap; 8.XII.1999 – 10.I.2000, 7♀ INB 3369853, 3369856, 3369859–63 ( INBC) ; 9.II–10.III.2000, 16♀ INB 3369926, 3369928–33, 3369941–8, 3369950 ( INBC) ; 10.III–6.IV.2000, 4♀ INB 3377241, 3377243–4, 3377246 ( INBC) , Malaise trap; Fila Catalina , Sector Catalina [10.33, ­85.25], 250 m , 8.I–8.II.2000, 3♀ INB 3369689–90, 3369896 ( INBC) , Malaise trap; Extremo E Campo de Aterrizaje , Sector Palo Verde [10.37, ­85.38], 0–50 m , 7.II–6.III.2000, 1♂ INB 3369919 ( INBC) , Malaise trap. HEREDIA. Santo Domingo , INBioparque, Santa Rosa, Bosque Seco Habitat [9.97, ­84.09], 1100 m , 8–29.II.2000, Zuniga, R., 1♀ INB 3370328 ( INBC) , Malaise trap; –. III.2000, 2♀ INB 3370298, 3369966 ( INBC) , Malaise trap. PUNTARENAS. Monteverde [10.483, ­84.833], 1400 m , 20.II.1987, Wood, D. M., 1♂ MEI 141523 ( CNC) ; Camino a Quijada del Diablo [8.89, ­82.76], 1500 m , 23.II.1997, Navarro, E., 1♀ INB 2501535 ( INBC) ; Coto Brus, Frontera con Panama , Sabalito , Estación Mellizas [8.89, ­82.77], 1400–1500 m , 28.XII.1995, 4♀ INB 3369962–4, 3369966 ( INBC) . SAN JOSE. Escaszu [9.917, ­84.133], 1179 m , 1–7.II.1988, FDP, 1♀ MEI 007978 ( EMUS) ; 7–14.II.1988, 1♀ MEI 007981 ( EMUS) ; 14–21.II.1988, 4♀ MEI 007969–71, 007976 ( EMUS) ; 16–23.II.1988, 1♀ MEI 007961 ( EMUS) ; 19.II.1988, 3♀ MEI 007974–5, 007977 ( EMUS) ; 4–7.III.1988, 1♀ MEI 007962 ( EMUS) ; 29.III–3.IV.1988, 2♀ MEI 007972–3 ( EMUS) ; 4–10.IV.1988, 1♀ MEI 007968 ( EMUS) ; 25–30.IV.1988, 8♀ ME I 007960, 007963–7, 007979–80 ( EMUS) ; 2–13.V.1988, 1♀ MEI 101167 ( EMUS) ; Escazú. [9.92, ­84.15] , 8.II.1987, Bohart, G. E., 1♀ MEI 007947 ( EMUS) ; 24–30.I.1988, FDP, 1♀ MEI 007940 ( EMUS) ; 1–7.II.1988, 6♀ MEI 007935, 007937, 007941–3, 007945, ( EMUS) ; 7–14.II.1988, 1♂ MEI 128821 ( INHS) , 3♀ 007936, 007950, 007951 ( EMUS) ; 14–21.II.1988, 1♂ MEI 007932 ( EMUS) ; 19.II.1988, 1♂ MEI 007924, 1♀ 007944 ( EMUS) ; 4–7.III.1988, 1♂ MEI 007926 ( EMUS) ; 8–18.III.1988, 1♂ MEI 007925, 3♀ 007953–4, 046190 ( EMEC) ; 21–27.III.1988, 2♂ MEI 007928–9 ( EMUS) ; 26.III.1988, 1♂ MEI 007922 ( EMUS) ; 29.III–3.IV.1988, 1♂ MEI 007923, 2♀ MEI 007952, 007955 ( EMUS) ; 1.IV–8.IV.1988, 2♂ MEI 007920–1, 2♀ 007948–9 ( EMUS) ; 19–24.IV.1988, 1♀ MEI 007931 ( EMUS) ; 25–30.IV.1988, 2♂ MEI 007927, 007930, 4♀ 007956–9 ( EMUS) ; 28.V.1988, 1♀ MEI 007934 ( EMUS) ; 11.VI.1989, 1♂ MEI 007919 ( EMUS) . MEXICO. JALISCO. Puerto Vallarta, Rio San Nicolas , Carr, Barra Novidad, Km 81 [19.2, ­104.69] , 27.I.1989, Ramírez, E., 1♂ MEI 106340 ( EBCC) ; 7.IV.1989, 1♂ MEI 050568 ( EBCC) . Sinaloa. 87 km S Culiacán [24.019, ­107.4], 165 m , 23.IV.1969, MEI, 3♂ MEI 040215, 040220–1 ( MEIC) , dry wash. YUCATAN. N Merida, Progreso Beach [20.967, ­89.617] , 29.VI.1966, 1♂ MEI 084498 ( SEMC) . NICARAGUA. MASAYA. Laguna de Apoyo [11.97, ­86.1] , 10–15.I.1992, van den Berghe, E., 1♂ MEI 039682 ( CMNH) , blacklight. VENEZUELA. ARAGUA. El Limon [9.93, ­67.02], 450 m , 17–18.II.1973, Rosales , C. J., 3♀ MEI 121561, 121569, 121601 ( MIZA) , Malaise trap; 19.II.1973, 1♀ MEI 121577 ( MIZA) ; 27.II.1973, 1♂ MEI 121613, 5♀ 121568, 121574, 121583, 121607, 121611 ( MIZA) ; 4–6.III.1973, 7♀ MEI 121571, 121580, 121599–600, 121602–3, 121605 ( MIZA) ; 12.III.1973, 1♀ MEI 121593 ( MIZA) ; 14.III.1973, 1♀ MEI 121587 ( MIZA) ; 15.III.1973, 3♀ MEI 121573, 121596, 121610 ( MIZA) ; 19.III.1973, 1♀ MEI 121564 ( MIZA) ; 21.III.1973, 1♀ MEI 121590 ( MIZA) ; 28.III.1973, 1♂ MEI 121612, 1♀ 121563 ( MIZA) ; 2.IV.1973, 1♀ MEI 121579 ( MIZA) ; 6.IV.1973, 1♀ MEI 121606 ( MIZA) ; 12.IV.1973, 2♀ MEI 121562, 121578 ( MIZA) ; 27.IV.1973, 1♀ MEI 121604 ( MIZA) ; 29.IV.1973, 1♂ MEI 121617 ( MIZA) ; La Victoria [10.231, ­67.345] , 11.II.1995, Marshall, S. A., 1♂ MEI 073579 ( DEBU) . CARABOBO. Bejuma [10.176, ­68.263] , 24.IV.1976, Michelangelli , C., Clavijo, J. A., 1♂ MEI 121616 ( MIZA) . Mé RIDA. 12 km SW Ejido, Rio Chama Valley [8.484, ­71.306], 1219 m , 17.II.1978, Heppner, J. B., 1♂ MEI 048182 ( USNM) , blacklight, thorn forest, dry.