Labeobarbus, in De Weirdt & Teugels, 2007

Vreven, Emmanuel J. W. M. N., Musschoot, Tobias, Snoeks, Jos & Schliewen, Ulrich K., 2016, The African hexaploid Torini (Cypriniformes: Cyprinidae): review of a tumultuous history, Zoological Journal of the Linnean Society 177 (2), pp. 231-305 : 302

AFRICAN LABEOBARBUS S.L. SPECIES: LIST OF EXAMINED (TYPE) SPECIMENS

acuticeps , Barbus Matthes, 1959 . Holotype: MRAC 130313. Paratypes: MRAC 130310 (1), 130311– 312 (2), 130314 (1). – acutirostris , Barbus brunelli Bini, 1940 . Neotype: RMNH 32870 (*). – aeneus , Cyprinus Burchell, 1822 . No types known. Synonyms: Barbus gilchristi Boulenger, 1911 . Holotype: BMNH 1909.12.8.1; Barbus mentalis Gilchrist & Thompson, 1913 . Holotype: SAIAB 134770 (ex SAM 9644). – altianalis , Barbus Boulenger, 1900 . Lectotype: BMNH 1906.9.6.13. Paralectotypes: BMNH 1906.9.6.14 (1), 1906.9.6.15 (1). Synonyms: Barbus eduardianus Boulenger, 1901 . Holotype: BMNH 1906.9.7.41; Barbus fergusonii Boulenger, 1901 . Syntypes: BMNH 1906.9.7.42–43 (2); Barbus kiogae Worthington, 1929 . Syntypes: BMNH 1929.1.24.105–108 (7 instead of 4); Barbus lobogenys Boulenger, 1906 . Syntypes: 1906.5.30.117–121 (2 instead of 5). – altipinnis , Varicorhinus Banister & Poll, 1973 . Holotype: MRAC 179729. Paratypes: BMNH 1972.10.2.1 (1), MRAC 179730 (1), 179731 (1), 179732-733 (2), 179734 (1), 179735 (1), IRSNB 643 (1). – ansorgii , Varicorhinus Boulenger, 1906 . Holotype: BMNH 1904.5.2.161. – aspius , Barbus Boulenger, 1912 . Syntypes: BMNH 1912.4.1.354 (1), MRAC 1536 (1). – axelrodi , Varicorhinus Getahun, Stiassny & Teugels, 2004 . Holotype: MRAC 91-68-P-1132. Paratypes (partim): MRAC 90-057-P-1297-1300 (4), 90-057-P-1314-1327 (14), 99-55-P-246-247 (2), 99-55-P-249 (1), 99-90-P-459- 461 (3). – batesii , Barbus Boulenger, 1903 . Holotype: BMNH 1902.11.12.128. – beso , Varicorhinus Rüppell, 1835 . Neotype: BMNH 1902.12.13.365. Specimens: BMNH 1968.7.24.18. – boulengeri , Labeobarbus (present paper). Lectotype: BMNH 1911.6.1.6. Paralectotypes (partim): BMNH 1911.6.1.7–10 (4), NMW 48865 (1), ZMB 18211 (1). – brauni , Varicorhinus Pellegrin, 1935 . Syntypes: MNHN 1935-0066 (1), MRAC 42933 (1). – brevicephalus , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 32880 (*). – brevispinis , Barbus ruspolii Holly, 1927 . Syntypes: NMW 7315 (1), 7316 (1). – bynni , Barbus Forsskål, 1775 . Neotype: BMNH 1907.12.2.1230. Synonyms: Barbus meneliki Pellegrin, 1905 . Holotype: MNHN 1905-0275; Barbus occidentalis Boulenger, 1911 . Holotype: BMNH 1909.3.3.14; Barbus seguensis Pellegrin, 1925 . Holotype: MNHN 1925- 0193. – cardozoi , Barbus Boulenger, 1912 . Syntypes (partim): BMNH 1912.4.1.343–348 (6), 1912.4.1.349– 352 (4), 1912.4.1.353 (1), MRAC 1528-29 (2), 1530-32 (3), ZMB 18810 (6). – caudovittatus , Barbus Boulenger, 1902 . Syntypes: BMNH 1901.12.26.26 (1), MRAC 1168 (1). Synonyms: Barbus euchilus Boulenger, 1920 . Holotype: BMNH 1919.7.24.7; Barbus miochilus Boulenger, 1920 . Syntypes: BMNH 1919.7.24.8-9 (2), MRAC 6992 (1); Varicorhinus stappersii Boulenger, 1917 . Syntypes: BMNH 1920.5.25.36-37 (2), MRAC 14197 (1), 14222 (2 instead of 1). – clarkeae , Varicorhinus Banister, 1984 . Holotype: MRAC 164456. Paratype: MRAC 164457 (1). – claudinae , Barbus De Vos & Thys van den Audenaerde, 1990 . Holotype: MRAC 86-01-P-501. Paratypes: MRAC 91755-756 (2), 92214 (1), 130310 (1), 85-44-P-141-144 (4), 85-44-P-269 (1), 85-44-P-281- 289 (9), 86-01-P-502-508 (7), 86-09-P-444-451 (8), 87- 11-P-1366-372 (7), 87-11-P-1373-384 (12). – codringtonii , Barbus Boulenger, 1908 . Holotype: BMNH 1908.11.6.23. Synonyms: Barbus altidorsalis Boulenger, 1908 . Holotype: BMNH 1908.11.6.26; Barbus chilotes Boulenger, 1908 . Syntypes: BMNH 1908.11.6.24–25 (2). – compiniei , Barynotus Sauvage, 1879 . Holotype: MNHN A-2845 (stuffed). Synonym: Barbus labiatomimus Pellegrin, 1914 . Holotype: MNHN 1886-0395. – crassibarbis , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 32890 (*). – dainellii , Barbus Bini, 1940 . Neotype: RMNH 32900 (*). – dartevellei , Barbus Poll, 1945 . Holotype: MRAC 47781. – dimidiatus , Varicorhinus Tweddle & Skelton, 1998 . Holotype: SAIAB 53080. Paratypes: SAIAB 53079 (15), SAIAB 53083 (21). – ensifer , Varicorhinus Boulenger, 1910 . Syntypes: BMNH 1910.11.28.134–143 (10), NMW 48864 (10), ZMB 18213 (8). – ensis , Barbus Boulenger, 1910 . Syntypes (partim): BMNH 1911.6.1.11–18 (8), NMW 54083 (4), ZMB 18217 (4). – ethiopicus , Barbus Zolezzi, 1939 . Holotype: Not seen. Specimens: BMNH 1971.7.12.1–3 (3). – fasolt , Barbus Pappenheim, 1914 . Holotype: ZMB 19061. – fimbriatus , Varicorhinus sandersi Holly, 1926 . Syntypes (partim): NMW 7224–7226 (3). – fritschii , Barbus Günther, 1874 . Syntypes: BMNH 1874.1.30.27– 31 (5). Synonyms: Capoeta atlantica Boulenger, 1902 . Syntypes: BMNH 1902.1.4.18–19 (2); Barbus paytonii Boulenger, 1911 . Syntypes: BMNH 1903.10.29.17–20 (7 instead of 4); Barbus riggenbachi Günther, 1902 . Syntypes: BMNH 1902.7.28.19 (1), 1902.7.28.20–21 (2); Barbus rothschildi Günther, 1901 . Syntypes: BMNH 1901.4.26.6–7 (2); Capoeta waldoi Boulenger, 1902 . Syntypes. BMNH 1902.1.4.16–17 (2). – gananensis , Barbus Vinciguerra, 1895 . Holotype: MSNG 17525 (*). Paratypes: MSNG 17331 (4) (*). – gestetneri , Barbus Banister & Bailey, 1979 . Holotype: BMNH 1976.10.12.98. Paratypes: BMNH 1976.10.12.86–97 (12). – girardi , Barbus Boulenger, 1910 . Syntypes (partim): BMNH 1911.6.1.31–36 (6), NMW-54137 (1), ZMB 18215 (1). – gorgorensis , Barbus intermedius Bini, 1940 . Neotype: RMNH 32910 (*). – gorguari , Barbus Rüppell, 1835 . Holotype: SMF 2586 (stuffed) (*). – gruveli , Barbus Pellegrin, 1911 . Holotype: MNHN 1911-0040 (*). – gulielmi , Barbus Boulenger, 1910 . Syntypes (partim): BMNH 1911.6.1.29–30 (2). – habereri , Barbus Steindachner, 1912 . Holotype: NMW 7274. – harterti , Barbus Günther, 1901 . Syntypes BMNH 1901.7.26.4–5 (2). – huloti , Barbus Banister, 1976 . Holotype: IRSNB 558. Paratypes: BMNH 1975.4.30.1 (1), IRSNB 563 (10). – humphri , Barbus Banister, 1976 . Holotype: IRSNB 559. Paratypes: BMNH 1975.4.30.2 (1), IRSNB 564 (10). – intermedius , Barbus Rüppell, 1835 . Holotype: SMF 6778 (stuffed) (*). Synonyms: Capoeta bingeri Pellegrin, 1905 . Holotype: MNHN 1905-0252; Barbus gudaricus Boulenger, 1906 . Syntypes: BMNH 1908.1.20.131– 132 (6 instead of 2), 1908.1.20.133 (1); Barbus leptosoma Boulenger, 1902 . Syntypes: BMNH 1902.12.13.300– 302 (3); Barbus macmillani Boulenger, 1906 . Syntypes: BMNH 1908.1.20.103–106 (4), 1937.4.20.68 (1) [a subsequently catalogued syntype (J Maclaine, pers. comm., 2015)]; Barbus plagiostomus Boulenger, 1902 . Syntypes: BMNH 1902.12.13.271–272 (2), BMNH 1902.12.13.273 (1). – iphthimostoma , Varicorhinus Banister & Poll, 1973 . Holotype: MRAC 179736. Paratypes: BMNH 1972.10.2.2 (1), IRSNB 644 (1), MRAC 179737 (1). – iturii , Barbus Holly, 1929 . Holotype: NMW (lost). – jaegeri , Varicorhinus Holly, 1930 . Holotype: NMW 13957. – johnstonii , Barbus Boulenger, 1907 . Holotype: BMNH 1897.6.9.280. Synonyms: Barbus globiceps Worthington, 1933 . Syntypes: BMNH 1932.11.15.275–282 (8); Barbus njassae Keilhack, 1908 . Syntypes: ZMB 18163 (1); 18164 (1); Varicorhinus nyasensis Worthington, 1933 . Syntypes: BMNH 1932.11.15.387–392 (6), 1932.11.15.393–395 (3). – jubae , Varicorhinus Banister, 1984 . Holotype: BMNH 1976.7.1.13. Paratypes: BMNH 1976.7.1.14–15 (2). – jubbi , Barbus Poll, 1967 . Holotype: Not seen. Paratypes (partim): MRAC 161065 (ex MD 2299) (1), 161066 (ex MD 6363) (1). – kimberleyensis , Barbus Gilchrist & Thompson, 1913 . Holotype: SAIAB 134771 (ex SAM 9645). – lagensis , Barynotus Günther, 1868 . Holotype: BMNH 1866.3.8.12. – latirostris , Barbus intermedius Keilhack, 1908 . Lectotype: ZMB 18174. Paralectotype: ZMB 34766. leleupanus , Varicorhinus Matthes, 1959 . Holotype: MRAC 92213. Paratypes: MRAC 92211 (1), 92212 (1), 130532–33 (2). – longidorsalis , Varicorhinus Pellegrin, 1935 . Holotype: MNHN 1935-0065. – longifilis , Barbus altianalis Pellegrin, 1935 . Lectotype: MNHN 1935-0150. Paralectotypes: 1935-0145 (1), 1935-0146 (1), 1935-0147 (1), 1935-0148 (1), 1935-0149 (1), 1935- 0151 (1), 1935-0152 (1). – longissimus , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 32930 (*). – lucius , Barbus Boulenger, 1910 . Syntypes: BMNH 1911.6.1.42–43 (2), NMW 54246 (1). – lufupensis , Varicorhinus Banister & Bailey, 1979 . Holotype: BMNH 1975.9.5.1. Paratype: BMNH 1975.9.5.2 (1). – macroceps , Barbus Fowler, 1936 . Holotype: ANSP 65759 (*). – macrolepidotus , Varicorhinus Pellegrin, 1928 . Syntypes: MNHN 1928-0011 (1), MRAC 19945 (1), 138767 (1; ex MNHN 1928-0012), NMB 3983 (1), 3985 (1), 3988 (1), 3989 (1). – macrolepis , Barbus Pfeffer, 1889 . Lectotype: ZMH H330 (ex 68 79). Paralectotypes: BMNH 1909.2.25.8 (ex ZMH)(1), ZMB 31672 (ex ZMH)(2), ZMH H331 (ex 68 79)(2), H332 (ex 380/7467)(4), H333 (ex 69 19)(1), H474 (ex 69 03)(1). – macrophtalmus , Barbus gorguarii Bini, 1940 . Neotype: RMNH 32940 (*). – malacanthus , Barbus Pappenheim, 1911 . Holotype: ZMB 18392. – marequensis , Barbus (Cheilobarbus) Smith, 1841 . Holotype: BMNH 1845.7.3.95 (stuffed). Synonyms: Barbus brucii Boulenger, 1907 . Holotype: BMNH 1907.3.15.34; Varicorhinus nasutus Gilchrist & Thompson, 1911 . Holotype: SAIAB 134736 (ex SAM 8801); Barbus oliphanti Keilhack, 1910 . Holotype: BMNH 1907.3.15.37; Barbus sector Boulenger, 1907 . Holotype: BMNH 1907.3.15.35; Barbus zambezensis Peters, 1852 . Syntypes: NMW 49730 (1); ZMB 3246 (2), 4744 (7). – mariae , Varicorhinus Holly, 1926 . Syntypes: NMW-7221 (1), 7222–7223 (2). – maroccana , Pterocapoëta Günther, 1902 . Syntypes: BMNH 1902.7.28.37–38 (2), 1902.7.28.39 (1). – matris , Barbus Holly, 1928 . Holotype: NMW 8000. – mawambi , Barbus Pappenheim, 1914 . Holotype: ZMB 19062. – mawambiensis , Barbus hindii Steindachner, 1911 . Syntypes: NMW 54177 (2), 54286 (3), 54287 (2), 54288 (2). – mbami , Barbus perplexicans Holly, 1927 . Holotype: NMW 7528. – megastoma , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 32950 (*). – micronema , Barbus Boulenger, 1904 . Syntypes: BMNH 1904.2.29.37– 38 (2). – mirabilis , Barbus Pappenheim, 1914 . Holotype: ZMB 19059. – mungoensis , Barbus Trewavas, 1974 . Holotype: BMNH 1973.5.14.163. Paratypes: BMNH 1973.5.14.164–182 (19). – nanningsi , Labeobarbus de Beaufort, 1933 . Holotype: ZMA 113010. – natalensis , Barbus de Castelnau, 1861 . Syntypes: lost. Synonyms: Barbus bowkeri Boulenger, 1902 . Syntypes: BMNH 1862.8.28.3–7 (5), 1874.3.5.1–2 (2), 1894.7.10.4 (1) [Note: BMNH 1862.8.28.3–7 non-conspecific syntypes of Enteromius gurneyi (Günther, 1868) ]; Barbus lobochilus Boulenger, 1911 . Holotype: BMNH 1908.12.28.96; Barbus mfongosi Gilchrist & Thompson, 1913 . Holotype: SAIAB 135057 (ex SAM 11392); Barbus robinsoni Gilchrist & Thompson, 1913 . Holotype: SAIAB 135055 (ex SAM 11371); Barbus zuluensis Gilchrist & Thompson, 1913 . Holotype: SAIAB 134939 (ex SAM 10745). – nedgia , Labeobarbus Rüppell, 1835 . Holotype: SMF 2619 (stuffed) (*). – nelspruitensis , Varicorhinus Gilchrist & Thompson, 1911 . Syntypes: SAIAB 134824 (ex SAM MB-F010518)(1), 135756 (ex SAM MB- F021698)(1). – nthuwa , Labeobarbus Tweddle & Skelton, 2008 . Holotype: SAIAB 39341. Paratypes: SAIAB 39293 (1), 40787 (6), 51928 (2), 79494 (2). – osseensis , Barbus Nagelkerke & Sibbing, 2000 . Holotype: RMNH 33721 (*). – oxyrhynchus , Barbus Pfeffer, 1889 . Lectotype: ZMH H339 (ex 69 07). Paralectotypes: ZMH H340 (ex 69 07)(7). Synonyms: Barbus babaulti Pellegrin, 1926 . Holotype: MNHN 1926-0285; Barbus hindii Boulenger, 1902 . Syntypes (partim): BMNH 1902.5.26.25–28 (4); Barbus (Labeobarbus) labiatus Boulenger, 1902 . Holotype: BMNH 1902.5.26.37; Barbus perplexicans Boulenger, 1902 . Syntypes: 1902.5.26.35-35 (2); Barbus tanensis Günther, 1894 . Syntypes (partim): BMNH 1893.12.2.24–29 (6), 1893.12.2.32–34 (3), BMNH 1893.12.2.37–39 (3). – pagenstecheri , Barbus Fischer, 1884 . Lectotype: ZMH H341 (ex ZMH 3851). Paralectotypes: ZMH H342 (ex ZMH 3850)(1). – parawaldroni , Barbus Lévêque, Thys van den Audenaerde & Traore, 1987 . Holotype: MRAC 73-10- P-2296. Paratypes: MRAC 73-10-P-2297 (1), 73-10-P- 2298-2299 (2), 73-5-P-1936 (1). – paucisquamata, Barbus altianalis Pellegrin, 1935 . Lectotype: MNHN 1935- 0076. Paralectotypes: MNHN 1935-0077 (1), 1935- 0078 (1), MRAC 42932 (1). Synonyms: Barbus altianalis lobogenysoides Pellegrin, 1935 . Holotype: MNHN 1935- 0154. – pellegrini , Varicorhinus Bertin & Estève, 1948 . Holotype: MNHN 1932-0181. – petitjeani , Barbus Daget, 1962 . Syntypes: MNHN 1959-0108 (3). – platydorsus , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 32970 (*). – platyrhinus , Barbus Boulenger, 1900 . Holotype: BMNH 1906.9.6.12. – platystomus , Varicorhinus Pappenheim, 1914 . Syntypes: ZMB 19051 (1), ZMB 34769 (1). – pojeri , Barbus Poll, 1944 . Holotype: IRSNB 71. Paratype: IRSNB 599 (1). – polylepis , Barbus Boulenger, 1907 . Holotype: BMNH 1907.3.15.33. – progenys , Barbus Boulenger, 1903 . Holotype: BMNH 1902.11.12.127. – pungweensis , Varicorhinus Jubb, 1959 . Holotype: SAIAB 120014 (ex AMG 850). Paratypes: SAIAB 120015 (ex AMG 851)(1), 120016 (ex AMG 852)(1), 120017 (ex AMG 853)(1), 120018 (ex AMG 854)(1). – reinii , Barbus Günther, 1874 . Syntypes (partim): BMNH 1874.1.30.22–24 (3). – rhinoceros , Barbus Copley, 1938 . Types?: BMNH 1936.12.22.35– 39. Synonyms: Barbus mariae Holly, 1929 . Lectotype: NMW 96552 (ex 6562)(designated by Banister, 1973). Paralectotypes: NMW 96553 (ex 6562)(1). – rhinophorus , Barbus Boulenger, 1910 . Syntypes: BMNH 1911.6.1.37– 38 (2). – robertsi , Varicorhinus Banister, 1984 . Holotype: BMNH 1983.3.30.20. Paratypes: BMNH 1983.3.30.21- 38 (18). – rocadasi , Barbus Boulenger, 1910 . Syntypes (partim): BMNH 1911.6.1.19–20 (2), 1911.6.1.21–25 (6 instead of 5), 1911.6.1.27–28 (2), NMW 13347–13352 (6), ZMB 18214 (6). – rosae , Barbus Boulenger, 1910 . Syntypes: BMNH 1911.6.1.39–41 (3), NMW 54500 (1), ZMB 18216 (1). – roylii , Barbus Boulenger, 1912 . Syntypes (partim): BMNH 1912.4.1.338–340 (3), 1912.4.1.341–342 (2), MRAC 1533 (1), 1534 (1), 1535 (1), NMW 54501 (1), ZMB 18809 (1). – ruandae , Varicorhinus Pappenheim, 1914 . Holotype: ZMB 19050. – ruasae , Barbus Pappenheim, 1914 . Lectotype: ZMB 19053. Paralectotype: ZMB 22652 (1). – ruwenzorii , Capoëta (Pterocapoëta) Pellegrin, 1909 . Syntypes: MNHN 1909-0583 (1), 1909-0584 (1), 1909-0585 (1). – sacratus , Barbus Daget, 1963 . Syntypes: MNHN 1959-0119 (4), 1959-0139 (1). – sandersi , Varicorhinus Boulenger, 1912 . Lectotype: BMNH 1912.4.1.333. Paralectotypes: BMNH 1912.4.1.334–336 (2 instead of 3), 1912.4.1.337 (1), MRAC 1525 (1), 1526 (1), 1527 (1), ZMB 18808 (1). – seeberi , Barbus Gilchrist & Thompson, 1913 . Syntypes: BMNH 1936.8.4.6 (1), SAIAB 134867 (ex SAM 10672)(2). Specimens: SAIAB 54133 (1), 54688 (1), 58362 (1), 58418 (1), 65536 (6). – semireticulatus , Varicorhinus Pellegrin, 1924 . Syntypes: MNHN 1924-0052 (2). – somereni , Barbus Boulenger, 1911 . Holotype: BMNH 1911.7.26.1. – stappersii , Barbus Boulenger, 1915 . Holotype: MRAC 14250. Synonyms: Barbus curtus Boulenger, 1915 . Holotype: MRAC 14172; Barbus moeruensis Pellegrin, 1922 . Holotype: MRAC 14765; Barbus oxycephalus Boulenger, 1915 . Holotype: MRAC 14233 (Note: not MRAC 14113 as in Lévêque & Daget, 1984; Eschmeyer, 2015). – steindachneri , Varicorhinus Boulenger, 1910 . Syntypes (partim): BMNH 1910.11.28.147–155 (9), 1911.6.1.1–5 (5), NMW 48867 (3), ZMB 18212 (3). – stenostoma , Varicorhinus Boulenger, 1910 . Holotype: BMNH 1910.11.28.145. – surkis , Barbus Rüppell, 1835 . Neotype: RMNH 32980 (*). – tornieri , Varicorhinus Steindachner, 1906 . Holotype: NMW 46053. – trachypterus , Barbus Boulenger, 1915 . Holotype: MRAC 11830. Synonym: Varicorhinus bredoi Poll, 1948 . Holotype: IRSNB 76. Paratypes: IRSNB 77 (2). – tropidolepis , Barbus Boulenger, 1900 . Lectotype: BMNH 1906.9.6.19. Paralectotypes: BMNH 1906.9.6.20–21, MRAC 327 (1), 328 (1), 329 (1). Synonym: Varicorhinus chapini Nichols & La Monte, 1950 . Holotype: AMNH 18785. – truttiformis , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 32990 (*). – tsanensis , Barbus Nagelkerke & Sibbing, 1997 . Holotype: RMNH 33000 (*). – upembensis , Varicorhinus Banister & Bailey, 1979 . Holotype: BMNH 1975.9.5.5. Paratypes: BMNH 1975.9.5.6–12 (7), 1975.9.5.13–17 (5). – urotaenia , Barbus Boulenger, 1913 . Syntypes: BMNH 1912.12.6.5 (1), MRAC 1791 (1), 1792 (2). – varicostoma , Varicorhinus Boulenger, 1910 . Holotype: BMNH 1910.11.28.146. – versluysii , Barbus Holly, 1929 . Syntypes: NMW 13954– 13956 (3). – werneri , Varicorhinus Holly, 1929 . Syntypes: NMW 13948 (1). – wittei , Varicorhinus Banister & Poll, 1973 . Holotype: MRAC 179738. Paratypes: BMNH 1972.10.2.3 (1), MRAC 179739–741 (3), 179742 (1), IRSNB 645 (1). – wurtzi , Barbus Pellegrin, 1908 . Holotype: MNHN 1908-0097. Specimens cf. wurtzi [undescribed species (see text)]: MRAC 1986-13-P- 114; MNHN 1959-0153 (2) (smallest of both: 127.5 mm SL), MNHN 1987-0689 (3), 1988-1955(3) (both largest specimens: 172.5 and 175.6 mm SL), 1991- 0519. – xyrocheilus , Varicorhinus Tweddle & Skelton, 1998 . Holotype: SAIAB 53082. Paratypes: SAIAB 34336 (11).