Banyallarga (Histricoverpa) sylvana, Prather Table Of Contents, 2004

Prather Table Of Contents, Aysha L., 2004, Revision of the Neotropical caddisfly genus Banyallarga (Trichoptera: Calamoceratidae), Zootaxa 435 (1), pp. 1-76 : 32-33

Banyallarga (Histricoverpa) sylvana , new species

Figs. 33–34

This species is known from Costa Rica and Nicaragua. Across this range, there is subtle variation in the shape of tergum X (Fig. 33A, B); however, I observed intermediate morphologies between the two extreme variants illustrated here and am confident that all are conspecific. The prominent dorsolateral lobe of tergum X (Fig. 33A, B) is the diagnostic character of this species, separating it from B. quincemil and acutiterga .

Adult. Forewing length 8.8–10.5 mm (n=15).

Head golden brown. Maxillary palps golden brown. Antenna twice forewing length, golden brown; each flagellomere pale basally and with pale strip of sensilla anteriorly. Dorsal pterothorax golden brown; ventrolateral thorax and legs golden brown, metathoracic tibia of male with long posterior setal fringe. Forewing uniform chestnut brown. Hind wing of male with basal semimembranous sleeve or pouch, enclosing brush of long setae; hind wing of female with short thick basal tuft of setae.

Male. Lateral margin of sternum IV deeply excavated (as in Fig. 16 ). Sternum VII with pointed anteromesal process. Tergum IX with anterior marginal ridge extending mesally to posterior margin, covered dorsally by semisclerotized membrane bearing dense fine pilosity (Fig. 33B); lateral ridge incomplete anteriorly; dorsal pleural setae fine and numerous (Fig. 33A). Preanal appendage longer than tergum X, slightly expanded apically and flexible, apical half covered with fine setae (Fig. 33B). Tergum X cleft posteromesally, separating roundly (Fig 33B 2) or acutely tapered (Fig. 33B 1) apicoventral projections; with paired rounded dorsolateral lobes (Fig. 33A, B); in lateral view with posteroventral projection (Fig. 33A). Inferior appendage simple, with long setae ventrally and laterally on coxopodite; harpago of uniform diameter along length, with fine setae basally and peglike setae apically (Fig. 33C). Phallus with horseshoe­shaped phallotremal sclerite, thornlike setae on endothecal membrane (Fig. 33D, E).

Female. Sternum VII with pointed anteromesal process. Sternum VIII anterior marginal ridge darkly sclerotized; fine setae covering posterior surface of sternite ( Fig. 34A ). Tergum IX with anterior marginal ridge not extending mesally, posterior margin not distinct from tergum X ( Fig. 34B ); sternum IX with anterior and posterior lobes darkly sclerotized and striate, with fine punctation posteriorly; anterolateral surface invaginated and rugose ( Fig. 34A ). Tergum X semimembranous, appendage rounded posteriorly ( Fig. 34B ); sternum X with semisclerotized plates bearing fine setae lateral to anal opening ( Fig 34A ). Vaginal apparatus anterior sclerite truncate anteriorly, posterolateral projections short; posterior sclerite ovoid anteriorly; dorsal connective membrane with regular pleating ( Fig. 34A ).

Holotype male: COSTA RICA: Alajuela: Reserva Forestal San Ramón, Río San Lorencito & tribs., 10°12'58"N 84°36'25"W, 980 m, 1–4.v.1990, Holzenthal & Blahnik ( UMSP000000068 ; UMSP).

Paratypes: COSTA RICA: Alajuela: unnamed river, Cerro Campana ca. 6 km (air) NW Dos Rios , 10°54'00"N 85°24'00"W, 640 m, 22–23.vii.1987, Holzenthal, Morse, & Clausen — 1 male ( NMNH) ; Parque Nacional Rincón de la Vieja, Quebrada Provisión, 10°46'08"N 85°16'52"W, 810 m, 4.iii.1986, Holzenthal & Fasth — 1 male ( UMSP) ; Reserva Forestal San Ramón, Río San Lorencito & tribs., 10°12'58"N 84°36'25"W, 980 m, 1–4.v.1990, Holzenthal & Blahnik — 2 males, 1 female ( UMSP) ; 2–4.vii.1986, Holzenthal, Heyn, & Armitage — 1 male, 1 female ( UMSP) ; 24–27.ii.1987, Chacón — 1 male ( UMSP) ; 30.iii–1.iv.1987, Holzenthal, Hamilton, & Heyn — 1 male, 1 female ( INBIO) ; Guanacaste: Río Los Ahogados, Río Los Ahogados , 11.3 km ENE Quebrada Grande, 10°51'54"N 85°25'23"W, 470 m, 26.vi.1986, Holzenthal, Heyn, & Armitage — 1 female ( UMSP) ; San José: Parque Nacional Braulio Carrillo , La Ventana, 1330 m, 5.viii.1990, 1 male ( INBIO) ; NICARAGUA: Jinotega: Peñas Blancas , 13°17'00"N 85°33'00"W, 1300 m, 25.vii.1997, Maes & Hernández — 1 male ( UMSP) ; Zelaya: Cerro Saslaya , 13°44'00"N 85°01'00"W, 700 m, 1.iv.1996, Maes & Hernández — 1 male, 1 female ( NMNH) .

Distribution. Costa Rica, Nicaragua.

Etymology. Sylvana, from the Latin Silvanus , meaning "god of woods," referring to the forests where this species was collected.

Figures

FIGURE 16. Banyallarga (Histricoverpa) acutiterga. Male (UMSP00000051): abdominal segments IV–X, lateral view. FIGURE 17. Banyallarga (Histricoverpa) acutiterga. Male (A, D, E, holotype; B, C, UMSP000000056): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view; E—phallotremal sclerites, dorsal view.

FIGURE 34. Banyallarga (Histricoverpa) sylvana. Female (UMSP000063348): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view.