Anhoraeomorphus ivelonanus (Franz) Jałoszyński, 2024

Jałoszyński, Paweł, 2024, The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 5505 (1), pp. 1-96 : 14-15

Anhoraeomorphus ivelonanus (Franz) , comb. n.

( Figs 28–35 )

Horaeomorphus ivelonanus Franz, 1986b: 171 .

Type material studied. Holotype (Madagascar): ♂, four labels: “Réserenat. III / Andranomalaza / Valeé d’Ivelona / X. 57 P. Soga” [white, printed], “INSTITUT / SCIENTIFIQUE / MADAGASCAR ” [bluish, printed], “ Horaeomorphus / ivelonanus m. / det. H.Franz ” [yellowish, handwritten and printed], “Typus” [red, handwritten] ( NHMW).

Revised diagnosis. Body ( Fig. 28 ) slender; head ( Fig. 29 ) slightly elongate, with vertex semicircular, in anterodorsal view its posterior margin evenly rounded; pronotum ( Fig. 28 ) subtrapezoidal, broadest at base, with two pairs of deep antebasal pits; antennomeres 1–7 distinctly elongate, 8–10 each about as long as broad; all femora similarly slender; metatibiae in males unmodified; aedeagus ( Figs 32–35 ) in ventral view broadest near middle, with apical region of dorsal wall weakly curved distoventrally, distolateral endophallic sclerites curved mesad and overlapping at middle, each with rounded apex, each paramere with three long apical setae.

Redescription. Body of male ( Fig. 28 ) light brown, tarsi and palps distinctly lighter, antennae slightly infuscate in distal halves, setae yellowish; BL 1.60 mm.

Head ( Figs 29–30 ) in anterodorsal view subtrapezoidal, broadest at eyes, HL 0.30 mm, HW 0.28 mm; temple in lateral view ( Fig. 30 ) slightly shorter than eye; vertex and frons confluent, together weakly convex and slightly elongate; posterior margin of vertex weakly and evenly rounded; each eye large and strongly convex, with small but distinct posteromedian emargination. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; frons and vertex densely covered with thick bristles directed posteriorly.Antenna ( Fig. 31 ) slender but short, slightly shorter than half BL, AnL 0.78 mm, antennomeres 1–7 each elongate, 8–10 each about as long as broad; 11 much shorter than 9 and 10 combined, slightly narrower than 10, nearly twice as long as broad.

Pronotum ( Fig. 28 ) subtrapezoidal, equally broad from base to middle; PL 0.40 mm, PW 0.33 mm. Anterior margin weakly arcuate, lateral margins slightly rounded in anterior half and strongly diverging posteriorly, in posterior half parallel; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.

Elytra ( Fig. 28 ) together oval, broadest near middle; EL 0.90 mm, EW 0.65 mm, EI 1.38; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.

Legs moderately long, slender, unmodified except slightly recurved tibiae.

Aedeagus ( Figs 32–35 ) stout, AeL 0.23 mm; in ventral view broadest near middle and weakly narrowing towards step-wise delimited apical region, which is subrectangular, transverse and shallowly emarginate; in lateral view apical region of dorsal plate weakly curved distoventrally; endophallus with slightly asymmetrical and elongate distolateral sclerites, symmetrical lateral groups of long and dense basal sclerites and asymmetrical weakly sclerotized median structures, distolateral sclerites strongly curved mesad and overlapping at middle, each with rounded apex; parameres thick, much shorter than median lobe, each paramere with three long apical setae.

Female. Unknown.

Distribution. Central Madagascar (Analamanga Region).

Remarks. According to Franz (1963 b), the word “Réserenat.” on the label is a misspelling of ‘Réserve nat’.

Anhoraeomorphus ivelonanus is externally slightly similar to A. grucheti , but it can be distinguished by the stronger elongate pronotum and distinctly elongate antennomeres 4–6 (about as long as broad in A. grucheti ). Another externally similar species is A. procerus . However, in A. procerus , the posterior margin of vertex is truncate and straight, whereas that in A. ivelonanus is rounded and posteriorly convex.

Figures

FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views.