Ancyropilus packsaddlensis, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 18-19

Ancyropilus packsaddlensis sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:E27871CE-9CEB-4109-B892-0C171352336E

( Figs 32 , 71 , 107 , 145 ; Map 1)

HOLOTYPE ♁: New South Wales: Packsaddle , 111 mi. N Broken Hill, 21-24 Nov. 1969 // Malaise trap H.E. Evans, R.W. Matthews ( ANIC) . PARATYPES (4): Queensland: Qld: 26.67° Sx 142.577°E 2.5km WNW Plevna Downs HS (PD3 M) 133m. 25 Nov 2008 - 17 Dec 2009. R. Mackenzie. Pitfall. Gidgee. 17317 (1 ♀, QM); Qld: 26.67°Sx142.577°E 2.5km WNW Plevna Downs HS (PD3 M) 131m. Gidgee. 15 Mar-3 Apr 2009. Malaise. R. Mackenzie. 17317 (1 ♀, 2, QM) .

Diagnosis. Pronotum angulate to almost rounded laterally, disc heavily punctate, glabrous areas in middle at base and either side of central impression; elytra dark with orange humeral, fasciate and apical maculations, fascia (with anterior and posterior margins approximately parallel) meeting at the suture, punctation lacking nodules, 8 th stria beginning within fascia, at least striae 3–9 reaching apical maculae; femora yellow basally, brown apically, tarsi with three ventral tarsal pads.

Description. Habitus: Fig. 145 . Total length: 6–7.1 mm (holotype 6.2 mm). Head: Cranium black to dark brown, submentum and gula bright orange, clypeus and supra-antennal elevation more reddish, anteclypeus orange, labrum, palpi and antennae orange-brown; eyes separated by 0.73–0.83 eye widths (holotype 0.73); vertex punctate, upper half of frons densely punctate, lower part punctate-rugulose until just before clypeus, clypeus with few punctations; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.6:1 (maxillae) and 2.6:1 (labium); antennae not reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter finer medially-directed setae. Prothorax: Pronotal disc and arch variously dark and light brown (pronotal arch and a medial discal stripe often lighter than remainder of disc), basal collar and venter orange to yellowish; pronotum 1.2–1.24 times longer than wide (holotype 1.22), sides angulate to almost rounded, middle wider than pronotal arch; subapical depression v-shaped, meeting shallow central impression; disc densely punctate, a thin medial glabrous stripe from base to central impression (difficult to view in paler specimens), area either side of central impression weakly tumescent, partially glabrous; short fine multi-directional setae (forming subtle swirls adjacent to glabrous areas) and long erect setae. Pterothorax: Ventrites orange to yellowish, with numerous pale, posteriorly- to posteromedially-directed setae; elytra dark brown with orange markings (each elytron with a squarish humeral macula, a very broad transverse fascia which meets at the suture and large apical macula); length to width ratio 2.59–2.72:1 (holotype 2.63); 8 th stria beginning within fascia (between its anterior margin and its middle), all striae reaching apical macula or striae 1, 2 and 10 not reaching apical macula, punctation lacking nodules, punctations posterior of fascia almost half the size of those anterior to fascia, epipleurae extending into apical maculae, interstriae with very fine short semi-erect setae and less frequent longer thicker erect setae, several setae often across interstical width; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad of CuA 1 crossvein small (but not completely absent). Legs: Basal two-thirds of profemora and basal three-quarters of meso- and metafemora yellow, all femora apically dark brown; tibiae and tarsi brown, ventral tarsal pads yellowish; meso- and metafemora slender, profemora slightly swollen. Abdomen: Ventrites orange to yellowish. Male genitalia: Tegmen ( Fig. 32 ) broadest near middle, narrowing before expanding again before apically acuminate parameroid lobes, dorsal sinus about one-quarter tegmen length, wide open, sides slightly convergent, internal lateral opening with short membranous lobes bearing a brush-like row of very short setae, ventral sinus about as deep, apodeme about one-quarter tegmen length; median lobe as in Fig. 71 ; pygidium as in Fig. 107 .

Variation. The paler brown area of the pronotum may be distinct or obscure.

Etymology. This species is named after the type locality of the male holotype, Packsaddle in north-western New South Wales.

Biology. Specimens were collected using pitfall, and Malaise, traps during the periods November-December and March-April. Traps at Plevna Downs were placed in Gidgee ( Acacia ) plant communities.

Distribution (Map 1). Ancyropilus packsaddlensis sp. nov. is known only from Packsaddle in north-western New South Wales and Plevna Downs in south-western Queensland.

Remarks. Similarities in body form, and general shape of the tegmen, suggest Ancyropilus packsaddlensis sp. nov. and A. emmotti sp. nov. are closely related. The decision to describe them as distinct species was based on correlations between consistant differences in the depth of the dorsal sinus of the tegmen and elytral patternation.

Figures

FIGURES 27–47. Tegmina, dorsal: (27) Ancyropilus brigalowae, paratype; (28) Ancyropilus emmotti, holotype; (29) Ancyropilus exossuarius, paratype; (30) Ancyropilus monteithi, holotype; (31) Ancyropilus noonbahensis, holotype; (32) Ancyropilus packsaddlensis, holotype; (33) Ancyropilus simplex, paratype; (34) Ancyropilus tricolor, paratype; (35) Ancyropilus hornensis, paratype; (36) Culcipternotus mareebensis, paratype; (37) Infectostriatus absentis, holotype; (38) Infectostriatus differens, holotype; (39) Monilonotum bunyense, paratype; (40) Monilonotum doddi; (41) Monilonotum pascoei; (42) Monilonotum sundholmi, paratype; (43–47) Notopilo, beswickensis species group—(43) Notopilo beswickensis, holotype; (44) Notopilo calicis, paratype; (45) Notopilo tompricensis, holotype; (46) Notopilo xanthoimprocerus, holotype; (47) Notopilo xanthoprolatus, holotype (scale bars: 0.5 mm).

FIGURES 66–101. Median lobes: (66) Ancyropilus brigalowae, paratype; (67) Ancyropilus emmotti, holotype; (68) Ancyropilus exossuarius, paratype; (69) Ancyropilus monteithi, holotype; (70) Ancyropilus noonbahensis, holotype; (71) Ancyropilus packsaddlensis, holotype; (72) Ancyropilus simplex, paratype; (73) Ancyropilus tricolor, holotype; (74) Ancyropilus hornensis, paratype; (75) Culcipternotus mareebensis, paratype; (76) Infectostriatus absentis, holotype; (77) Infectostriatus differens, holotype; (78) Monilonotum bunyense, paratype; (79) Monilonotum pascoei; (80) Monilonotum sundholmi, paratype; (81–85) Notopilo, beswickensis species group—(81) Notopilo beswickensis, holotype; (82) Notopilo calicis, paratype; (83) Notopilo tompricensis, holotype; (84) Notopilo xanthoimprocerus, paratype; (85) Notopilo xanthoprolatus, holotype; (86–88) Notopilo, cambageicola species group—(86) Notopilo cambageicola, paratype; (87) Notopilo interfabulatus, paratype; (88) Notopilo tanybasilaris, paratype; (89) Notopilo congruus (congruus species group); (90) Notopilo reduncus (reduncus species group); (91–93) Notopilo, variipes species group—(91) Notopilo brevistriatus, paratype; (92) Notopilo gerstmeieri, holotype; (93) Notopilo variipes; (94–96) Notopilo, unplaced to species group—(94) Notopilo elstoni, holotype; (95) Notopilo katherinensis, holotype; (96) Notopilo obesus, paratype; (97) Platynotum bulli, paratype; (98) Platynotum culgoense, holotype; (99) Platynotum femorale; (100) Platynotum foveosetosa, holotype; (101) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 138–149. Habitus: (138) Ancyropilus brigalowae, paratype; (139) Ancyropilus emmotti, holotype; (140) Ancyropilus exossuarius, paratype; (141) Ancyropilus hornensis, paratype; (142) Ancyropilus labris, holotype; (143) Ancyropilus monteithi, paratype; (144) Ancyropilus noonbahensis, holotype; (145) Ancyropilus packsaddlensis, holotype; (146) Ancyropilus simplex, paratype; (147) Ancyropilus tricolor, paratype; (148) Culcipternotus mareebensis, holotype; (149) Infectostriatus absentis, holotype (scale bars: 1.0 mm).