Anabarhynchus moretonensis Ferguson

Ferguson, David J., Lambkin, Christine L. & Yeates, David K., 2014, Eight new species of Australian stiletto flies in the genus Anabarhynchus Macquart (Diptera: Therevidae) from South East Queensland, Zootaxa 3802 (4), pp. 553-582 : 572-575

Anabarhynchus moretonensis Ferguson sp. n.

( Figs 17 , 18 , 19 )

Type material. Holotype: Male. AUSTRALIA: Queensland: 7km E. Dunwich, North Stradbroke Island, QLD; 27°30'S 153°27'E; 120m, G. Daniels, C.J. Burwell; ( GDCB _18189) (AM_K:257751) (AM). Condition: Micro pinned dorsally onto pith block; many occipital macrosetae lost.

Paratypes: 3 males and 2 females. AUSTRALIA: Queensland: 3♂, 2♀ same collection data as holotype; ( GDCB _18186) (AM_K:257745), ( GDCB _18187) (AM_K:257747); ( GDCB _18188) (AM_K:257762); ( GDCB _18190) (AM_K:257777); ( GDCB _18191) (AM_K:257819), (AM).

Diagnosis. Frons slightly raised with reflective grey pubescence; male frons width 3.3–3.8 × ocellus width, female 4.1–4.6 × ocellus width; pair of grey vittae on scutum; katepisternum without pile; prosternal furrow with pile; fore femur with 1 pd; hind femur, 1 av macroseta. Female sternite 8 with enlarged ‘posterior lobe’.

Description. Male. Body length: 7 mm. Wing length: 5.5 mm. ( Figs 17–19 ). Head. Integument black. Frons slightly raised, width 3.4 x ocellus width; ocellar tubercle flat; antennal base positioned low on frons; face and lower frons slightly protruding; parafacial and frons with reflective grey pubescence; two rows of black, long, erect frontal setae, mostly on upper frons; lower frons setae equal length of scape. Scape length 3 × pedicel length; scape and pedicel grey; f1 dark grey with several short black setae dorsally on the basal third; f2 and f3 black, combined length half that of f1. Occiput convex with grey pubescence; occipital macrosetae pale, in several rows, 45–48 each side; postocciput to gena grey, with long pale pile. Palp pale yellow with pale hair-like setae; labellum dark grey; prementum setae pale. Thorax. Integument black; scutal chaetotaxy black (pairs): np, 4; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with a pair of grey vittae either side of dark grey and brown mid-line stripe; indistinct brown marks outlining raised areas pre and post-transverse suture; short black appressed setae sparsely distributed. Katepisternum without pile; prosternal furrow with pile; pleura grey; coxae grey with long pile admixed with black macrosetae. Wing. Hyaline with grey infuscate, brown veins, and stigma brown; costal setae beyond humeral crossvein biserially arranged. Haltere. Pedicel stem yellowish grey; knob dirty white. Legs. Fore femur with 1 pd; hind femur 1 av macroseta; all femora dark grey, covered with grey pubescence and appressed pale pile on dorsal surfaces admixed with short black setae, ventral pile erect; tibiae and tarsi pale brown, apically darker. Abdomen. Integument black; tergites covered with thick pubescence that appears bright grey viewed anteriorly and black when viewed posteriorly; sternites blackish with grey pubescence. Terminalia. Paratype genitalia (GDCB_18187) (AM_K:257747): Epandrium ( Fig. 19 A): length approximately 3/4 its width (measured along mid-line and between widest points when viewed dorsally); pale setae over posterior half, admixed with several black setae on posterolateral edge. Gonocoxite ( Fig. 19 B): darkly sclerotised, appearing rounded when viewed ventrally, slightly longer than wide; broad lobe-shaped outer gonocoxal process with pale macrosetae admixed with a few black on the posterolateral margin; a patch of dense of pale pile on a raised projection each side of the ventromedial plane. Gonocoxite connected anteriorly by a robust, darkly sclerotised hypandrium, medially with a thinly sclerotised ‘window’, and by a short length along the ventromedial plane. Gonocoxal apodeme darkly sclerotised; short, extending just beyond anterior edge of gonocoxite. Inner gonocoxal process curved medioventrally, with broad flange on inner edge; many short black and pale setae apically, admixed with a few longer weak pale setae. Gonostylus long and narrow, directed mediodorsally, with dense long pale pile on dorsobasal and inner edges. Ventral lobe triangular with sparse pale setae on ventral edge. Aedeagus ( Figs 19 C, D): distiphallus narrow and ventrally directed; parameral sheath triangular, darkly sclerotised, curved dorsomedially apically; ventral apodeme anteriorly broad and flat with the apical margin slightly bilobed; lateral ejaculatory apodeme broad, band-like; ejaculatory apodeme long and darkly sclerotised.

Variation. Males, body length 6.5–8.0 mm; frons width 3.3–3.8 × ocellus width; occipital macrosetae: 45–47 each side; wing length: 6.0– 6.5 mm;

Female, body length 7.5 mm; frons brown grey; frons width 4.1-4.6 × ocellus width; occipital macrosetae 46–48 each side; wing length: 6.0– 6.5 mm; abdominal anterior bands viewed posteriorly appear matte black, viewed anteriorly matte brownish grey; tergite 2 and 3 with whitish apical bands. Paratype female (GDCB_18191) (AM_K:257819): Sternite 8 ( Fig. 19 F): 1.5 × longer than wide (measured along mid-line and between widest points when viewed dorsally); anterior margin flat, with triangular indentation medially; large thinly sclerotised oval area middle of posterior half with long thin black setae sparsely distributed anteriorly. Posterior with enlarged ‘posterior lobe’ covered with evenly distributed weak long setae. Furca ( Fig. 19 E) sarcophagus shaped, darkly sclerotised, posterior frame thin, tapered with slightly bilobed apex; medial frame strong with poorly sclerotised panels; anterior beam thin, supporting a large, darkly sclerotised arterial ventral lobe with posterolaterally directed projections that reach the frame.

Etymology. The specific epithet ‘ moretonensis ’ is derived from the geographical location of Moreton Bay that is formed in part by Stradbroke Island, where the type specimens were collected.

Comments. Known from specimens collected in March and April on North Stradbroke Island, south-east Queensland.

Anabarhynchus moretonensis sp. n. keys to A. albosetosus Lyneborg in couplet 24 in Lyneborg (2001) and can be separated from A. albosetosus which has a generally grey pubescent scutum with indistinct longitudinal vittae; frons width 3 x ocellus width (in the male); fore femur without pd macrosetae; inner gonocoxal process with a reduced flange on the sub-apical inner edge; and gonocoxal connection partly fused along the ventromedial plane. Anabarhynchus moretonensis sp. n. scutum has a pair of distinct grey vittae either side of a dark grey and brown medial stripe; frons width 3.3–3.8 × ocellus width (in the male) fore femur with 1 pd macroseta; inner gonocoxal process with a broad flange along inner edge, and gonocoxite not fused but weakly connected by thin sclerotised membrane along the ventromedial plane.

Anabarhynchus moretonensis sp. n. appears to be most closely aligned to species to Lyneborg’s niveus speciesgroup.

Figures

FIGURE 17. Anabarhynchus moretonensis sp. n., holotype male, (GDCB _ 18189) (AM _ K: 257751), lateral view. Scale line = 2 mm.

FIGURE 18. Anabarhynchus moretonensis sp. n., holotype male, (GDCB _ 18189) (AM _ K: 257751) oblique view. Scale line = 2 mm.

FIGURE 19. Anabarhynchus moretonensis sp. n., paratype male terminalia (GDCB _ 18187) (AM _ K: 257747); A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view; paratype female (GDCB _ 18191) (AM _ K: 257819): E, furca; F, female sternite 8, ventral surface. Scale line = 0.5 mm.