Amplirhagada uwinsensis, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 261-263

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
DOI	https://doi.org/10.5281/zenodo.5239045
persistent identifier	https://treatment.plazi.org/id/4CCC73A5-13FC-4FD9-A525-22DEFF33ADB3
taxon LSID	lsid:zoobank.org:act:4CCC73A5-13FC-4FD9-A525-22DEFF33ADB3
treatment provided by	Felipe
scientific name	Amplirhagada uwinsensis
status	sp. nov.

Treatment

Amplirhagada uwinsensis View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia, Kimberley, Bonaparte Archipelago, Hanover Bay , Uwins Island , 15°15'32"S 124°46'08"E; KIS-3–106. Vine thicket and large fig trees on upper slopes, piles of sandstone rocks, under large slabs. (leg. V. Kessner, 09 June 2006) GoogleMaps .

Type material. Holotype WAM S34621 View Materials (Pl. 2.7) . Paratypes AMS C463715 (15 preserved specimens, as holotype), WAM S36991 View Materials (35 preserved specimens, as holotype), AMS C463716 (15 shells, 15°15'25"S 124°48'04"E), WAM S37441 View Materials (30 shells, 15°15'25"S 124°48'04"E) GoogleMaps .

Additional, non-type material. WAM S36987 View Materials –90, WAM S37440 View Materials , WAM S37442 View Materials –4, AMS C463717 (Uwins Island).

Etymology. In reference to Uwins Island, where this species occurs.

Shell ( Fig. 69A–C View Figure 69 , Pl. 2.7–9). Semi-globose with medium high spire; solid. Periphery angulate; upper and basal sectors of whorls rounded. Umbilicus completely concealed by columellar reflection. Background colour light reddish brown; sub-sutural band diffuse to well defined, light brown to chestnut brown, varies greatly in thickness and contrast; mid-whorl band diffuse, brownish to chestnut brown, thin, varies greatly in contrast and thickness, visible on last whorl(s) only; bands may blend into each other across whorl surface; ventral colour yellowish-greenish brown to ochre; outer lip colour blending into chestnut brown; inner lip blending into dark ochre to chestnut brown. Protoconch c. 2.7 mm in diameter, comprising 1.7 whorls, with fine, indistinct axial lirae. Teleoconch with marked growth lines, becoming oblique towards base of shell. Angle of aperture 45°; outer lip rounded, slightly expanded, not reflected, basal node of lip weak, palatal node absent. Parietal wall of inner lip absent.

Pallial morphology. Pallial cavity short, extending ½ to 3 ⁄ 4 whorls; mantle pigmentation consists of indistinctive, sparsely distributed brown spots. Kidney extending about half or more than half of pallial cavity.

Genital morphology ( Figs. 67–68 View Figure 67 View Figure 68 ). Penis rather straight, more or less of same length as anterior part of oviduct. Vas deferens coils once before entering penis. Penial retractor muscle very short to stubby. Penial verge very short (barely visible), slender to spatulate, with pointed tip. Penial wall pustules comparatively large, elongated, arranged in rows on median and basal portion of penial chamber, forming densely packed, narrow longitudinal pilasters at apical end of penial chamber. Main stimulatory pilaster welldifferentiated, large, cone-shaped, covered with flattened pustules, comprising anterior to median portion of penial chamber. Two additional pilasters are formed that are covered by flattened pustules or smooth narrow ridges. Vas deferens entering penial sheath in upper third. Vagina moderately wide, tubular to posteriorly inflated; inner vaginal wall with weakly developed longitudinal pilasters. Spermathecal duct wide, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising less than half to about half of anterior part of oviduct, more or less straight. Spermoviduct of same length as anterior part of oviduct or longer. Talon embedded in albumen gland close to anterior end of albumen gland.

Radular morphology ( Fig. 69E–F View Figure 69 ). Rectangular. Tooth formula C+14–16+4+22. In average 149±16.0 rows of teeth, 29.5±1.0 rows per mm (n = 2). Central teeth with sharply pointed, triangular mesocones, shorter than base of teeth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of teeth; ectocones and endocones vestigial. Marginal teeth with elongate mesocones; ectocones smaller and narrower than mesocones; endocones shorter than ectocones.

Comparative remarks. One of the few species with large, dome-shaped shells. Differs from A. tricenaria and A. regia by conspicuously dark brown outer lip colour and by huge, cone-shaped main pilaster, presence of additional pilasters and characteristically elongated pustules of inner penial wall.