Adelopomorpha tuberculata, Will & Guéorguiev, 2021

Will, Kipling & Gueorguiev, Borislav, 2021, Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini), ZooKeys 1044, pp. 375-425 : 375

Adelopomorpha tuberculata sp. nov. Figs 28 , 38 , 41 , 53

Material examined.

Holotype: New Caledonia • ♂; " NEW CALEDONIA Mt. Do 0.5km below summit, 28Jan2004 G.B.Monteith "// " QM berleseate 1111 21°45'Sx 166°00'E, rainforest, 950 m, sieved litter" Holotype pinned, with genitalia in a separate microvial. Source collection QM, deposited MNHN. Type locality as given on label.

Paratypes: New Caledonia • ♂; " NEW CALEDONIA 11420 21°45' Sx 166°00'E, Mt. Do summit, 1000m, 22 Nov 2003, G. Monteith, Pyrethrum, trees & logs." [QM] • ♂; " NEW CALEDONIA Mt. Dzumac, 800-1000m, 23 May 1984 G. Monteith & D. Cook " [QM] .

Diagnosis.

This species is nearly indistinguishable externally from A. glabra except for the lack of the depressed lateral 1/3 of the pronotum present in A. glabra (Fig. 43 ), and its smaller size. Also, it is smaller than A. tethys , whose male has setae on ventrite 6. See diagnosis above. The aedeagus is decisively different (Fig. 28 ), with a relatively long and moderately, narrowly rounded tip.

Description.

Habitus. Very small sized, BL: 4.90 mm and BW: 2.45 mm (range, BL: 4.75-5.00 mm, BW: 2.45-2.60 mm), with rather oval and convex body (Fig. 41 ). Color and luster. Head dark reddish brown; pronotum, elytra and ventral surface of body black, pronotum marginally diaphanous reddish black; legs deep reddish brown. Deep reddish black medially at the very apical edge of ventrite 6; antennae and palpi brown. Integument moderately dull, with slight, diffused spectral iridescence dorsally and ventrally. Microsculpture and punctation. Dorsal surface of head and pronotum with prominent isodiametric meshes; elytra with irregular mesh of more or less transversely stretched sculpticells; ventral surface with scarcely-visible sculpticells on ventrite 6 and prosternum, otherwise sculpticells mostly not apparent. Head lacking macropunctation or wrinkles on vertex; head, pronotum and elytra without micropunctation. All abdominal ventrites very shallowly wrinkled to nearly smooth. Head. Slightly more than 1/4 as wide as pronotum (Suppl. material 2: Table S1). Eye moderate sized, somewhat protruded, EyW/HW: 1.31. Frontoclypeal sutures not evident. Mentum tooth without paramedial border. Thorax. Pronotum 1 2/3 wider than long (PW/PL: 1.67 (range, 1.71-1.76)), with sides evenly, very shallowly rounded from anterior angles to posterior ones; maximal width in posterior 1/5; width at apex ~ 1/3 that of widest point (PW/PA: 2.88 (range 2.82-3.00)). Posterior angles ~ right angled, broadly rounded. Mesosternum with single, very prominent, medial tubercle. Metepisternum 1 1/2 × wider than long, with lateral margin straight, coadunation with epipleuron along entire length. Elytra. Broad, as wide as long (EL/EW: 1.00 (range, 1.00-1.02)). Basal margin distinct, forming small, sharp tooth at shoulder, ended medially at level of parascutellar punctures, joined to parascutellar puncture fovea by a short, shallow striole. Apical sinuation not evident. Female genitalia. Female specimens unknown. Male genitalia. Median lobe of aedeagus in lateral view long, almost straight (Fig. 38 ); lobe in dorsal view with apical lamella moderately long, rounded, triangular and oriented to right (Fig. 28 ); sclerotized portion of endophallus visible in repose, with two sclerites.

Etymology.

The specific epithet Adelopomorpha tuberculata draws attention to the relatively large, median mesosternal tubercle in these beetles. It is treated as an adjective in the nominative singular.

Figures

Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima.

Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus.

Figures 41, 42. Dorsal habitus of 41 Adelopomorpha tuberculata 42 holotype of A. glabra Heller, with labels. Images of A. glabra by O. Jaeger, reproduced with permission from MTKD.

Figures 52, 53. Maps of localities of New Caledonian Oodini.

Figures 43, 44. Forebody, left dorsolateral view of 43 Adelopomorpha glabra Heller 44 Adelopomorpha glabra group male from Gelima.