Actenoncus wallacei, Tian, Ming-Yi & Deuve, Thierry, 2009

Tian, Ming-Yi & Deuve, Thierry, 2009, A review of the genus Actenoncus Chaudoir (Coleoptera: Caraboidea: Orthogoniini), Zootaxa 2135, pp. 57-64 : 60-62

Actenoncus wallacei , new species

( Fig. 2 )

Holotype. female, labeled “Homut/Wallace” only, in Bates’ Collection, MNHN. The collecting data of the unique specimen is missing but we presume that it was collected by Homut and Wallace from Indonesia.

Diagnosis. Medium-sized, dark brown or brownish, right mandible without median tooth, labrum straight at frontal margin, head with small punctures at frons and vertex, pronotum smooth, impunctate, elytral striae shallow, punctures large and connected one another.

Length: 15.5 mm; width: 6.3 mm.

Head and disc of pronotum dark brown, elytra, lateral expanded margins of pronotum, antennae and legs brownish. Habitus as in Fig. 2 .

Surface very shiny, head with sparse, small punctures only near eyes, frons and vertex; smooth and glabrous; pronotum impunctate, smooth and glabrous. Microsculptural meshes densely isodiametric.

Head as long as wide; eyes large, extraordinary prominent, neck constricted, labrum straight at frontal margin; frontal impressions deep, very wide, but short, extending only to the level of fore margins of eyes; clypeus bisetose, base distinctly raised, and with a furrow in median portion; mandibles stout, right mandible without median tooth; palpi subcylindrical, slightly dilated and stout; maxillary palpomere 3 as long as 4, both glabrous except palpomere 3 with two or three short setae at apex; labial palpomere 2 slightly longer than 3; labial palpomere 2 bisetose at subapex in inner margin; ligula narrow, bisetose; mentum without median tooth, asetose, submentum with a pair of setae; palpiger unisetose; antennae short and stout, only reaching the base of elytra; antennomeres 1 and 4 subequal, slightly longer than 3, and nearly twice as long as 2.

Pronotum strongly transverse, PW/PL=1.67, disc strongly convex; markedly expanded at sides, widest at about middle, lateral expanded margins rather narrow, evenly widened, and distinctly reflexed; fore and hind angles rounded, basal margin distinctly produced backwards in median portion; both transversal impressions well marked, basal foveae wide and deep.

Elytra elongate ovate, EL/EW=1.5, convex; base unbordered, widest a little behind middle, slightly paralleled at sides; apex broadly truncate, inner angle nearly rectangular, pointed; strial punctures large, wellmarked and striate (connected each other, especially in apical half, almost striate near apex), intervals flat, equal in width at middle, impunctate, except a row of tiny punctures at basal part of interval 6; interval 3 without setiferous pore.

Prosternal process unbordered at apex. Hind tarsomere 4 bilobed at apex. Abdominal ventrite VII complete at apical margin, bisetose on either side.

Male. Unknown.

Remarks. This new species differs from A. punctatus by the following morphological features: (1) pronotum impunctate; (2) front of labrum nearly straight (while emarginated in A. punctatus ); (3) elytral striae deep, evidently punctate and connected one another (striae shallow, with punctures larger but isolated in A. punctatus ); (4) pronotum with lateral expanded margins narrower, quite even, and reflexed (wider, uneven and flat in A. punctatus ); and (5) antennae stouter, only extending to base of elytra (but at 1/4 of basal elytra in A. punctatus ). Compared with A. foersteri , this new species is broader in form, pronotum impunctate, elytral striae largely punctate (but smaller in A. foersteri ), and right mandible without median tooth (evidently toothed in A. foersteri ).

Etymology. The specific name of this new species is dedicated to Alfred R. Wallace, a well-known British traveler and naturalist in the 19th century.

Distribution. Indonesia, detailed locality unknown.

Figures

FIGURES 1 – 4. Habiti of Actenoncus. 1. A. ater, male, 2. A. wallacei, n. sp., female, holotype, 3. A. punctatus, male, holotype, 4. A. foersteri, male. scale bar: 10.0 mm.