Cryobius Poecilus ventricosus Haptoderus Feronia spadicea Feronia pumilio Pseudorthomus Feronia amaroides Orthomus Pseudocryobius Feronia nivalis Cryobius Abaxodes Feronia abaxoides Abax Orites Platysma negligens Pyreneorites Feronia pusilla Orites Euhaptoderus Carabus unctulatus Haptoderus Iberoderus Feronia nemoralis Cryobiopterus Argutor brevicornis Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico Bousquet, Yves ZooKeys 2012 2012-11-28 245 1 1722 F22C5E07-941C-545F-ABA3-AC09C1FB1E3D Chaudoir, 1838 Chaudoir 1838 Insecta Carabidae Hypherpes GBIF Animalia (Cryobius) Coleoptera 542 543 Arthropoda subGenus Cryobius   CryobiusChaudoir, 1838: 11. Type species:  Poecilus ventricosusEschscholtz, 1823 by original designation. Etymology (original). From the Greek cryos(cold) and bios(life), alluding to the cold habitat where these species live [masculine].  HaptoderusChaudoir, 1838: 10. Type species:  Feronia spadiceaDejean, 1828 (=  Feronia pumilioDejean, 1828) by original designation. Synonymy established by Bousquet (1999: 183). Etymology (original). From the Greek hapto(fasten to, by extension to apply) and dere(neck, by extension pronotum) [masculine].  PseudorthomusChaudoir, 1838: 12, 19. Type species:  Feronia amaroidesDejean, 1828 by original designation. Synonymy established by Bousquet (1999: 183). Etymology (original). From the Greek pseudos(fallacy, lie) and the generic name  Orthomus[masculine].  PseudocryobiusMotschulsky, 1850a: ix. Type species:  Feronia nivalisSahlberg, 1844 designated by Bousquet (1984a: 4). Synonymy established by Poppius (1906b: 7). Etymology (original). From the Greek pseudos(fallacy, lie) and the generic name  Cryobius[ q.v.] [masculine].  AbaxodesGistel, 1857: 31. Type species:  Feronia abaxoidesDejean, 1828 designated by Strand (1917: 76). Etymology. From the generic name  Abax[ q.v.] and the Greek - odes(likeness) [masculine].  OritesSchaum, 1858: 442, 463. Type species:  Platysma negligensSturm, 1824 by monotypy. Synonymy established by Schatzmayr (1942: 77). Etymology. From the Greek oreites(mountaineer) [masculine].  PyreneoritesJeannel, 1937b: 11, 16. Type species:  Feronia pusillaDejean, 1828 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the geographical name Pyrenees and the generic name  Orites[ q.v.] [masculine].  EuhaptoderusJeanne, 1969: 33. Type species:  Carabus unctulatusDuftschmid, 1812 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the Greek eu(agreeable, original, primitive) and the generic name  Haptoderus[ q.v.] [masculine].  IberoderusJeanne, 1969: 34. Type species:  Feronia nemoralisGraells, 1851 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the geographic name Iberia (the Iberian Peninsula) and the Greek dere(neck, by extension pronotum) [masculine].  CryobiopterusO. Berlov, 1997: 36. Type species:  Argutor brevicornisKirby, 1837 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the Greek cryos(cold), bios(life), and pteron(wing, by extension elytron) [masculine].  Diversity. About 115 species in the arctic, subarctic, and boreal regions of North America (23 species) and the Palaearctic Region (about 100 species). Several species are Holarctic.   Identification. Ball (1966a) revised the North American species which he arrayed in three groups. Since then, two new species have been described, one (  Pterostichus haftorni) by Lindroth (1969a) which has been subsequently synonymized, the other one (  Pterostichus woodi) by Ball and Currie (1997), and several species related to  Pterostichus planushave been downgraded to subspecies of  Pterostichus bryantiby Ball and Currie (1997). Species identifications are difficult in part because of the important intraspecific variation observed in most species.  Taxonomic Note. Lorenz (2005: 275) listed  ParahaptoderusJeanne as a junior synonym of  Cryobius. However, the taxon, which is composed of three European species of which one extends to Turkey, is probably not closely related to  Cryobius(see Bousquet 1999: 187).