An integrated taxonomic revision of Ctonoxylon (Coleoptera, Curculionidae, Scolytinae) reveals new Malagasy species originating from multiple recent colonisations of the island Jordal, Bjarte H. ZooKeys 2024 2024-05-28 1203 95 130 Jordal, 2024 Jordal 2024 Insecta Curculionidae Ctonoxylon Animalia Coleoptera Arthropoda species tuberculatum sp. nov.   Figs 30, 33, 36   Type material.    Holotype, male: Madagascar, Diana prov., Montagne d’Ambre[GIS: - 12.54, 49.17],  1000 malt., ex  Ficusbranch,  03. 11. 2019, B. Jordal, leg. [  ZMUB].  Allotype[  ZMUB] and paratype[  NHMW]: same data as holotype.   Diagnosis. Two teeth along the anterior margin of pronotum separated by more than width of a tooth; scattered interstrial setae separated within uniseriate rows by more than their length; posterior margin of elytra and declivital interstriae with sharp tubercles, largest tubercles on interstriae 1 and 3.   Description.  Male.Body length 3.0– 3.1 mm, 2.1–2.2 × as long as broad; colour brown. Fronsflattened from upper level of eyes to epistoma, surface shrivelled; vestiture of fine short setae. Eyes divided, each part separated by a little more than size of upper half. Antennal funiculus 7 - segmented; club setose, sutures and septum barely indicated. Pronotumcoarsely asperate on anterior three quarters, two front teeth separated by little more than width of a tooth, located behind the margin. Anterior lateral margin of prothorax at middle with prominent eye scraper; a sharp carina running from eye scraper to near procoxa; propleural pit absent. Scutellar shieldwider than long, oval. Elytralstriae impressed, punctures irregular and small; interstriae rounded, densely micropunctate, vestiture consisting of uniseriate rows of slightly curved bristle-like setae and fine dense ground vestiture along the suture; elytral suture straight; elytral apex slightly emarginated, along the posterior margin and at each declivital interstriae with small sharp tubercles, at interstriae 1 and 3 tubercles 2–3 × larger. Metanepisternumwith mixed short bifid and longer bristle-like setae; metaventrite with simple hair-like setae; these sclerites on its anterior third partly glabrous and with small bifid setae, and with a swollen trace of mesotibia’s resting position. Femaleas in male, except anterior pair of teeth more closely placed and located along the anterior margin of the pronotum, and surface of the frons smooth.   Etymology. The Latin adjective tuberculatusin its neuter form, reflecting the tubercles on declivital interstriae which are more prominent than in related species.   Distribution and biology. Only known from the holotypelocality in Madagascarwhere it was collected from very thick bark of a fallen  Ficusbranch ( Moraceae), 7 cmin diameter. Two pairs were collected at the early stage of tunnel construction, including a mating niche with a short egg tunnel.   Comments. Previous records of  C. flavescensfrom Madagascarare most likely  C. tuberculatumas these two species are very similar. This could also be the case for records of  C. montanum(recorded as  C. longipilum), in which the female mainly differs by the more closely set anterior teeth on the pronotum, and the stouter body. These two species are therefore removed from the list of Malagasy species. 4869223359 ex Ficus branch 2019-11-03 ZMUB B. Jordal Madagascar Diana prov. 1000 Montagne d'Ambre Montagne d'Ambre 1 holotype 4869223366 ex Ficus branch 2019-11-03 ZMUB, NHMW B. Jordal Madagascar Diana prov. 1000 Diana prov. Montagne d'Ambre 2 allotype