New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London Darby, Michael European Journal of Taxonomy 2019 2019-04-04 512 1 50 8Y6PB Darby, 2019 [619,785,1479,1506] Insecta Ptiliidae Niptella GBIF Animalia Coleoptera 24 25 Arthropoda species gutta gen. et sp. nov.   urn:lsid:zoobank.org:act: B6E16C2B-3B88-418C-9536-6E235936248C  Fig. 12    Diagnosis Determined by the form of the mesoventral keel.    Etymology From the Latin ‘  gutta’, meaning ‘drop’, and referring to the shape of the mesoventral keel.    Type material    Holotype MALAYSIA-BORNEO• ♀; Sarawak, 4 thdivision, Gunung Mulu National Park;  1930 ma.s.l.; May–  Aug. 1978; P.M. Hammondand J.E. Marshallleg., BM 1978-49; BMNH.    Paratypes MALAYSIA-BORNEO• ♂; same data as for holotype; BMNH•  2 specs; same data as for preceding;  1800-1900 ma.s.l.; litter; BMNH•  1 spec.(mounted disarticulated on the slide); Sarawak, 4 thdivision, Gunung Mulu National Park;  May–Aug. 1978; P.M. Hammondand J.E. Marshallleg., BM 1978-49; litter below comp. 4; BMNH•  1 spec.; Sarawak, Mulu;  13 Mar. 1978; N.M. Collinsleg.; pitfalls in alluvial forest; BMNH.    Description SIZE. Habitus ( Fig. 12A), length 0.65 mm. COLOUR. Yellowish brown shining, closely covered with fine pale pubescence. HEAD. With row of postulate setae behind the eyes ( Fig. 12F). Antennae very pale yellow almost translucent, antennomeres III–XI, length 0.28 mm. Eyes prominent, distance across eyes 0.24 mm. Mentum and submentum ( Fig. 12D). PRONOTUM. Length 0.19 mm, width 0.30 mm( Fig. 12F).   Fig. 12.  Niptella gutta gen. et sp. nov. A. Habitus. B. Spermatheca. C. Aedeagus. D. Mentum and submentum ×860. E. Pygidium. F. Pronotum ×540. G. Meso- and metaventra ×470. H. Mesoventral keel ×1060. ELYTRA. Length 0.39 mm, width 0.35 mm. truncate. VENTRUM. Proventrum wide before contiguous procoxae, reticulate, with scattered setae. Mesoventrum ( Fig. 12G), collar not extending onto sharply toothed humeri, keel in shape of a peardrop stopping short of collar and narrowing to a point between the mesocoxae ( Fig. 12H). Metaventrum ( Fig. 12G), metacoxae separated by half the width of the sclerite. ABDOMEN. Pygidium ( Fig. 12E). FEMALE GENITALIA. Spermatheca globular, very small and difficult to detect ( Fig. 12B). MALE GENITALIA. Aedeagus large ( Fig. 12G). 2234828129 1978-08 BMNH P. M. Hammond & J. E. Marshall MALAYSIA-BORNEO 1930 Gunung Mulu National Park 24 25 BM 1978-49 1 1 Sarawak holotype 5006575310 1978-08 BMNH P. M. Hammond & J. E. Marshall MALAYSIA-BORNEO 1930 MALAYSIA-BORNEO 25 26 1 1 Sarawak paratype 2234828194 1978-08 BMNH P. M. Hammond & J. E. Marshall Malaysia 1850 Gunung Mulu National Park 25 26 2 Sarawak paratype 2234828128 1978-08 1978-08 1978-05 BMNH P. M. Hammond & J. E. Marshall Malaysia Gunung Mulu National Park May-Aug. 25 26 BM 1978-49 1 Sarawak paratype 2234828178 1978-03-13 BMNH N. M. Collins Malaysia Mulu 25 26 1 Sarawak paratype