Review of the genus Togoda Melichar, 1906 (Hemiptera, Auchenorrhyncha, Fulgoroidea, Tropiduchidae), with descriptions of three new species from tropical Africa and Madagascar Gnezdilov, Vladimir M. Zoosystema 2025 2025-09-30 47 23 473 488 Gnezdilov 2025 [334,507,1171,1198] Insecta Tropiduchidae Togoda Animalia Hemiptera 9 480 Arthropoda species minima sp. nov.  ( Figs 5A, B; 6E, F; 10)  urn:lsid:zoobank.org:act: B19D2836-E909-4258-8652-D09F1F525DFD     TYPE  MATERIAL. —  Holotype.  Mozambique• ♂; “Museum Paris / Moçambique / Vallée du Pungouí / Guengére / G. Vasse1906” [grey blue] // “Février” [white] // “Museum Paris / MNHN(EH) / 16572” [white]; MNHN.    Paratype.  Mozambique• ♀; “Museum Paris / Mozambique / Vila Pery / P. Lesne1928” [grey blue] // “Mars” [white] // “Museum Paris / MNHN(EH) / 16571” [white]; MNHN.   DIAGNOSIS. — Fore wings small, with 1/5 of its length behind apex of clavus in lateral view. Metopial proboscis without notch frontally. Phallobase long, nearly as long as aedeagus, asymmetric.  ETYMOLOGY. — Species name referring to small fore wings.   DISTRIBUTION. — Central Mozambique(Nhamatanda District).   DESCRIPTION  Structure ( Figs 5A, B; 6E, F) Coryphe weakly transverse, 1.5 times as wide as long medially, without carinae, anterior margin straight, posterior margin nearly straight ( Fig. 5A). Metope, pro- and mesonotum densely covered with pustules ( Fig. 6F). Coryphe-metope margin keel-shaped ( Figs 5B; 6E). Metopial proboscis without notch frontally ( Fig. 6F). Post- and anteclypeus with fine median carina medially which is running from the middle of postclypeus throughout anteclypeus. Eyes large, diameter of each eye nearly as wide as coryphe. Median part of pronotum with a groove medially. Fore wings rather short, nearly oval, surpassing by 1/5 of its length beyond abdominal apex in male and reaching abdominal apex in female ( Fig. 5B). Basal cell narrowly oval. Radius and median running from basal cell by one short stem or from one point and firstly furcating near to the cell in basal fifth of wing. Forewing vein branching sequence: R 5, all branches terminating at costa not reaching wing margin; r-m 4; M 10, firstly furcating near to basal cell, M 1and M 2again furcating after wing middle; im 12; m-cua 4-6; CuA 2, furcating near wing middle; icua 3. Pcu joint A 1near to wing middle. Clavus long, 4/5 of wing length. Legs not flattened neither foliated. Hind tibia with nine spines apically. First metatarsomere with two latero-apical and 7-9 intermediate spines arranged in an arc apically.  Coloration ( Fig. 5A, B) General coloration yellowish brown. Head laterally light yellow below antennae, with wide black stripe above scapi. Coryphe, pronotum, and mesonotum yellowish brown. Each paranotal lobe of pronotum with large dark brown to black area proximally. Mesonotum sometimes with dark brown lateral parts. Metope brown to dark brown, with reddish lateral margins and light yellow pustules. Postclypeus yellowish brown to dark brown, with pale median carina. Anteclypeus light yellow. Rostrum light yellow, with black apex. Scapi and pedicels brown. Each paranotal lobe of pronotum with large dark brown to black area proximally. Mesonotum sometimes with dark brown lateral parts. Forewing yellowish brown to dark brown, with basal subcostal field brown to dark brown, some cells of corium sometimes dark brown to black, and veins reddish brown. Basal processes of forewing precostal margins dark brown to black. Hind wings dark brown. Fore femora light yellowish brown to dark brown. Fore tibiae with three dark brown bands. Fore tarsi dark brown. Apices of leg spines black. Claws dark brown. Thorax and abdomen below light yellow. Abdominal sternite III light yellow. Abdominal sternites IV-VI yellowish brown. Female sternite VII with black laterally hind margin. Gonoplac margins dark brown, with black teeth. Gonocoxae VIII dark brown, with black hind margins proximally. Male genital block yellowish brown.   FIG. 6. — Head of  Togodaspp.: A, B,  T. africana Melichar, 1906, male, Togo (Bismarckburg), head, lateral view ( A), metopial proboscis, frontal view ( B); C, D,  T. excisa n. sp., paratype, head, lateral view ( C), metopial proboscis, frontal view ( D); E, F,  T. minima n. sp., paratype, head, lateral view ( E); metopial proboscis, frontal view ( F); H, G,  T. carinata n. sp., metopial proboscis, frontal view ( G); holotype, head, lateral view ( H). Scale bar: 0.5 mm.  Male terminalia ( Fig. 10) Pygofer wide in dorsal half, with strongly convex hind margins (in lateral view) ( Fig. 10A). Anal tube elongate, twice as long as wide medially, narrowing to truncate apex (in dorsal view) ( Fig. 10C).   FIG. 7. —  Togoda africana Melichar, 1906, Togo (Bismarckburg), male: A, right hind wing; B, rostrum. Abbreviations: A1, first anal vein; A2, second anal vein; CuA, cubitus anterior; CuP, cubitus posterior; M, median vein; Pcu, postcubitus; R, radial vein. Scale bars: A, 1 mm; B, 0.5 mm. Apex of anal tube slightly turned downwards (in lateral view) ( Fig. 10B). Anal column short and wide. Penis wide, horse-shoe shaped (in lateral view) ( Fig. 10D, E). Phallobase long, nearly as long as aedeagus, assymetric, with ventral lobe shifted to the left side and narrowing apically. Aedeagus with a single hookshaped subapical process turned downwards ( Fig. 10D, E, a) on right side. Phallobase with long and wide plate-shaped process, with a short tooth-shaped projection apically, turned upwards on right side ( Fig. 10D, e) and narrow pointed apically and turned upwards process on left side ( Fig. 10D, E, f). Style with nearly straight hind margin and large apical tooth ( Fig. 10F, G, at). Connective with elongate large cup ( Fig. 10D).  Female terminalia Anal tube elongate, narrowing to truncate apex.  Measurement Total length. Male: 6.0 mm. Female: 6.5 mm. 5835980302 TYPE, MATERIAL, MNHN G. Vasse Mozambique Mozambique 9 480 1 1 holotype 5835980307 MNHN P. Lesne Mozambique Mozambique 9 480 1 1 paratype