Lenapterus Lyperopherus vermiculosus Pterostichus Galapterus Pterostichus galae Pterostichus Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico Bousquet, Yves ZooKeys 2012 2012-11-28 245 1 1722 7A6060E5-4A9A-5A86-A410-858CE0B65227 Berlov, 1996 Berlov 1996 Insecta Carabidae Euferonia GBIF Animalia (Lenapterus) Coleoptera 518 519 Arthropoda subGenus Lenapterus   LenapterusO. Berlov, 1996: 11. Type species:  Lyperopherus vermiculosus Menetries, 1851 by original designation. Etymology (original). From the name of the river Lena and the Greek pteron(wing, by extension elytron) or the first two syllables of the generic name  Pterostichus[ q.v.] [masculine].  GalapterusO. Berlov and Plutenko, 1997: 47. Type species:  Pterostichus galae Farkacand Plutenko, 1996 by original designation. Synonymy established by Sundukov (2005: 803). Etymology. From the first name of Plutenko'swife and the Greek pteron(wing, by extension elytron) or the first two syllables of the generic name  Pterostichus[ q.v.] [masculine].  Diversity. Eleven species in the arctic, subarctic, and boreal regions of North America (four species) and Asia (ten species, one of them extending to European Russia). Three species are Holarctic.  Identification. Lindroth (1966) reviewed the North American species. Budarin (1976) reviewed the world fauna, including two species (  Pterostichus mirus Tschitscherineand  Pterostichus rugosusGebler) that belong, however, to the subgenus  MetallophilusChaudoir. Recently Sundukov (2005) revised all known species and provided a key for their identification.  Taxonomic Note. This subgenus has been known for sometimes under the name  LyperopherusMotschulsky (e.g., Ball 1960b: 127; Lindroth 1966: 525).