Meskea dyspteraria Grote, 1877 Meskea horror Dyar, 1913 Meskea dyspteraria Taxonomic revision of the monotypic genera of Siculodini (Lepidoptera, Thyridoidea, Thyrididae, Siculodinae) Queiroz-Santos, Luziany Casagrande, Mirna Martins Mielke, Olaf Hermann Hendrik Zootaxa 2021 2021-01-28 4920 2 235 253 3ZZRR Grote, 1877 ♀ Grote 1877 [151,540,906,933] Insecta Thyrididae Meskea Animalia Lepidoptera 9 244 Arthropoda species dyspteraria  ( Figs 4, 8, 12, 15)    Meskea dyspterariaGrote, 1877. The Canadian Entomologist, 9: 115; 1 female[holo] type, Bastrop, Texas, [ USA], Heiligbrodt leg., Meske collection, Albany [ USA].— Hampson, 1897. Proceedings of the Zoological Society of London, 1897(1): 632, fig. 24 (male d).— Dalla Torre, 1914. Lepidopterorum Catalogus 20: 43.— Gaede, 1936, inSeitz. Die Gross - Schmetterlinge des Amerikanischen Faunengebietes. Die amerikanischen Spinner und Schwärmer 6: 1211, fig. 175h (d).— Whalley, 1964. Annals and Magazine of Natural History, (13) 7(74): 122.— Whalley & Heppner, 1995, Thyridoidea, inBecker et al.Checklist: Part 2, Hyblaeoidea–Pyraloidea–Tortricoidea, inHeppner (ed.). Atlas of Neotropical Lepidoptera 3: 31.   Meskea horror Dyar, 1913. Proceedings of the United StatesNational Museum, 44: 317; [holo] type 1 male, Jalapa, Mexico, V-1908, R. M̹ller leg., N° 13370, USNM.— Gaede, 1936, inSeitz. Die Gross -Schmetterlinge des Amerikanischen Faunengebietes.Die amerikanischen Spinner und Schwärmer 6: 1211.— Whalley & Heppner, 1995, Thyridoidea, inBecker et al.Checklist: Part 2, Hyblaeoidea–Pyraloidea–Tortricoidea, inHeppner (ed.). Atlas of Neotropical Lepidoptera 3: 31.— Day et al., 1997. Entomophaga, 42(3): 393. Syn. nov.  Taxonomic history.  Meskea dyspteraria was described without illustration, based on a female [holo] typefrom Bastrop, Texas, USA.    Meskea horror was described without illustration, based on a male [holo]type from Jalapa, Mexico, May, 1908, R. M̹ller leg, nº 13370 and four paratypes: one male from Córdoba, Mexico, W. Schaus leg. and three females from Cuernavaca, [ Morelos], Mexico, June and July, 1906, W. Schaus leg., USNM.  Taxonomic comments. Dyar (1913)noted that  M. horroris distinguished from  M. dyspterariaby the apex of the hindwing being more tapered and the outer margin more emarginate. After examining the [holo]type of  M. dyspteraria, the [holo]type and the paratypesof  M. horror, genitalia, wing morphology and geographic distribution of males and females, we concluded that the hindwing differences described by Dyar (1913)represent variations of the same species. Thus,  M. horroris considered a new junior synonym ( syn. nov.) of  M. dyspteraria.    Typematerial.  Meskea dyspteraria: Thefemale [holo] typewas found in the USNM, with the author’s original labels, which correspond with the information provided in the original description. [Holo] typehaving the following labels: /  Meskea dyspteraria ♀Grote Type/ Tex / 230 / Collection O. Meske / TypeN° 339 U.S.N.M. /. Mistakenly, Whalley (1964)mentioned that the [holo] typeis a male deposited in the NHMUK.   Meskea horror: Themale [holo]type and the four paratypeswere found in the USNM, with the author’s original labels corresponding to the original description. [Holo]type with the following labels: / Jalapa Mex / May 08 / R Muller Collector / 1723 /  Meskea horrorType Dyar / Type N° 13370 U.S.N.M. / Ƌ genitalia Slide, 18 Oct. ’32 C.H. #2 / Genitalia Slide By ME Ƌ USNM110, 112 /, and the following labels are added: / HOLOTYPE/ HOLOTYPE  Meskea horror Dyar, 1913/. The four paratypesare labeled similarly to the [holo]type. The paratypefrom Córdoba, Mexicolisted as male, is actually a female.   Redescription. As for genus.   FIGURE 6. Male genitalia of  Iza nubeculafrom Brazil, Bahia, Camacan (Reserva Serra Bonita) (MVOB 145120). A) ventral view. B) lateral view. C) inner view of valva. D) dorsal view of tegumen and uncus. E) dorsal view of aedeagus. F) lateral view of aedeagus. G) ventral view of aedeagus. Scale bar 1mm.   FIGURE 7. Male genitalia of  Loxiorhiza cervinalisfrom Brazil, Mato Grosso do Sul, Miranda (Bodoquena mountain) (CEIOC 003). A) ventral view. B) lateral view. C) inner view of valva. D) dorsal view of tegumen and uncus. E) dorsal view of aedeagus. F) lateral view of aedeagus. Scale bar 1mm.   FIGURE 8. Male genitalia of  Meskea dyspterariafrom Costa Rica, Guanacaste (Santa Rosa National Park) (NHMUK 010921587) (valva and fultura inferior distorted due to preparation technique). A) ventral view. B) dorsal view of tegumen and uncus. C) lateral view of aedeagus. Scale bar 1mm.   FIGURES 9–12. Female genitalia of  Hepialodes,  Iza,  Loxiorhizaand  Meskeaspecies in lateral view. 9.  H. folliculafrom Brazil, Goiás, Campinaçu (Barra Mansa farm) (DZ 39.216). 10.  I. nubeculafrom Brazil, S„o Paulo, Salesópolis (Boraceia) (DZ 39.288). 11.  L. cervinalisfrom Bolivia, Santa Cruz, Buenavista (NHMUK 010921618). 12.  M. dyspterariafrom El Salvador, San Salvador (MSFR). Scale bar 1mm.  Geographic distribution( Fig. 15). USA: Arizona: Pima, Baboquivari Peak Wilderness; Alabama: Mobile; Mississippi: Harrison, Handsboro; Hanckock, Diamondhead; Florida: Washington, Chipley, Rock Hill Prsv.; Liberty, Torreya State Park; Leon, Tallahassee; Escambia, Pensacola, Miami-Dade, Baía de Biscayne, Paradise Key; Texas: Brown, Lake Brownwood State Park, San Jacinto, Coldspring, San Houston National Forest; Kimble, Junction; Burnet, Shovel Mountain; Montgomery, Conroe; Bastrop, Bastrop; Harris, Crosby; Chambers, Wallisville, Lake Charlotte; Uvalde, ConCan, Rio Frio; Bexar, San Antonio; Bexar, Ebony Hill Res Station (EHRS); San Patricio, Sinton, Welder National Wildlife Refuge; Hidalgo, Santa Ana National Wildlife Refuge; Pharr, Bentsen Rio Grande Valley State Park; Mercedes; Cameron, Laguna Atascosa National Wildlife Refuge, Sabal Palm Sanctuary, Brownsville. MEXICO: Sinaloa: Venadio; Durango; San Luis Potosí: Tamazunchale; Yucatán: Cansahcab; Veracruz:  Jalapa; Cardel; Veracruz; Córdoba; Orizaba; Presidio; Catemaco; Colima: Comala; Morelos: Cuernavaca; Estado de México: Zacualpan; Puebla: Tehuacán; Chiapas: La Polka, S. of Tonala; Mapastepec. GUA-TEMALA:  Retalhuleu: San Sebastian;  Guatemala:  Ciudad de Guatemala, Río Platanos. EL SALVADOR: San Salvador. COSTA RICA: Guanacaste: Santa RosaNational Park; Area de ConservaciónGuanacaste; Puntarenas: Aranjuez; San José; Cartago: Cachi. PANAMA: Panamá: La Cabima. VENEZUELA: Guárico: Hato Masaguaral, 45 kmS Calabozo. PERU: Lima: San Isidro.  Natural history. Immature induces galls in  Sida acuta Burman, 1768and  Sida rhombifolia Linnaeus, 1753(Malvaceae). Females oviposit on the abaxial side of the host plant leaf. Larvae form a gall on the stem which increases with larval size. An opening is made at the base to allow adult emergence. Larval development lasts approximately 192 days ( Day et al. 1997). The species occurs in forests and open areas, and adults were caught throughout the year other than December.   FIGURES 13–14. Signa of  Hepialodesand  Loxiorhizaspecies. 13.  H. folliculafrom Brazil, Goiás, Campinaçu (Barra Mansa farm) (DZ 39.216). Scale bar 1mm. 14.  L. cervinalisfrom Bolivia, Santa Cruz, Buenavista (NHMUK 010921618). Scale bar 0,5mm.  Examined material(103 Ƌ, 176 ♀total).   USA:  Arizona: Pima, Baboquivari Peak Wilderness, 1 ♀, 2 Ƌ,  1 –15.VII.1923, 1 Ƌ,  15 –30.VII.1923, 1 Ƌ,  15 –30.VIII.1923, 2 ♀,  15 –30.VI.1924, 1 ♀,  15 –30.VII.1924, 3 ♀,  15 –30.VIII.1924, C. Poling leg. ( USNM),  1 ♀,  1 –15.VIII.1923, 1 ♀,  1 –15.VII.1924, C. Poling leg., 1 ♀,  16.VI.1967, J. H. Hessel leg. ( AMNH);   Alabama:  Mobile, 1 ♀,  5.V.1921, C. Dukes leg., 1 Ƌ,  20.III.1922, 1 ♀,  25.IV.1923( USNM);   Mississippi: Harrison,  Handsboro, 1 ♀,  30.IV.1967, R. & B. Taylor leg. ( AMNH);  Hanckock,  Diamondhead, 1 ♀,  10.V.1991, H. Xeroobien leg. ( USNM);   Florida: Washington,  Chipley, Rock Hill Prsv., 1 ♀,  28.IV.2017, J. Hayden leg. ( MGCL);  Liberty, Torreya State Park, 1 ♀,  16.V.1968, H. V. Weems Jr. leg., 1 ♀,  15.V.1970, H. Greenbaum leg., 1 ♀,  21 –24.V.1987, H. O. Baggett leg., 1 ♀,  22.V.1987, Charles Stevens leg. ( MGCL);  Leon,  Tallahassee, 1 ♀, NHMUK 10921563( NHMUK);  Escambia,  Pensacola, 1 ♀,  13.V.1941, Shirley Hills leg. ( MGCL),  Miami-Dade, Baía de Biscayne, 1 ♀( AMNH),  Paradise Key, 1 ♀,  1.III.1910, 1 ♀,  III.1910, 1 ♀,  II.1927( USNM);   Texas: 2 ♀, 4 Ƌ,  13.IV.1924, 2 ♀,  21.IV.1924, 7 ♀, 1 Ƌ,  28.IV.1924, 1 ♀,  29.IV.1924, L. J. Bottimer leg., 1 ♀, H. C. Hanson leg., 1 ♀, Friedrich leg. ( MfN), 1 Ƌ (USNM);  Brown, Lake Brownwood State Park, 1 ♀,  8.V.1964, A. & M. E. Blanchard leg. ( USNM);  San Jacinto,  Coldspring, San Houston National Forest, 1 ♀,  8.V.1988, E. C. Knudson leg., 1 ♀, 1 Ƌ,  2.V.1990, Linwood C. Dow leg. ( MGCL);  Kimble,  Junction, 4 ♀,  20.V.1966, A. & M. E. Blanchard leg. ( USNM);  Burnet, Shovel Mountain, 1 ♀( USNM);  Montgomery,  Conroe, 1 ♀,  11.V.1969, A & M. E. Blanchard leg. ( USNM);  Bastrop,  Bastrop, 1 ♀([ Holo] typeof  Meskea dyspteraria); Harris,  Crosby, 1 Ƌ,  4 –10.V.1919( AMNH),  4 ♀, 5 Ƌ,  4 –10.V.1919, 1 ♀,  14.IV.1919, H. C. Hanson leg. ( USNM);  Chambers,  Wallisville, 1 Ƌ,  12.V.1919, ( AMNH), 3 Ƌ,  9.V.1919, 1 Ƌ,  9 –19.V.1919, C. P. Heinrich leg., 3 Ƌ, 1 ♀,  12.V.1919, 1 ♀, 3 Ƌ,  12 –15.V.1919, 2 Ƌ,  12 –19.V.1919, A. Busck leg. ( NMHN),  Lake Charlotte, 1 Ƌ,  11.IV.1923, 1 Ƌ,  17.IV.1923, 1 ♀, 1 Ƌ,  19.IV.1923, 1 ♀,  25.IV.1923( AMNH),  1 ♀, 6 Ƌ,  17.IV.1923, 3 ♀,  18.IV.1923, 1 Ƌ,  19.IV.1923, 1 ♀, 3 Ƌ,  20.IV.1923, 6 ♀, 1 Ƌ,  23.IV.1923, 1 ♀, 1 Ƌ,  25.IV.1923, 2 ♀, 1 Ƌ,  26.IV.1923, L. J. Bottimer leg. ( NMHN);  Uvalde,  ConCan, 1 ♀,  1.V.1992, 1 ♀,  2.V.1992, C. Bordelon leg., 1 ♀,  1 –2.V.1992, E. C. Knudson leg., 1 ♀, 1 Ƌ,  6 –7.V.2005, Bordelon& Knudson leg. ( MGCL),  Rio Frio, 1 Ƌ,  30.IV –2.V.1992, Ron Leuschner leg., 1 ♀,  9.V.1994, Kurt Leuschner leg. ( USNM);  Bexar, 2 Ƌ,  28.IV.1928, H. B. Parks leg. ( USNM),   San Antonio, 1 ♀,  16.V.1976, E. C. Knudson leg., 1 Ƌ,  17.V.1977, William W.& Nadine M. leg. ( MGCL);  Bexar, Camp Bullis Roadx Loop1604, 1 ♀,  1.IV.1977, William W.& Nadine M. leg. ( MGCL);  Bexar, Ebony Hill Res Station( EHRS), 1 Ƌ,  25.IV.1985, Roy O. Kendall& C. A. Kendall leg. ( MGCL);  San Patricio,  Sinton, Welder National Wildlife Refuge, 3 ♀,  20.IV.1963, 1 ♀,  12.V.1963, 1 ♀,  22.IV.1964, 1 ♀,  19.V.1965, A. & M. E. Blanchard leg. ( USNM);  Hidalgo, Santa Ana National Wildlife Refuge, 1 ♀, 1 Ƌ,  7.IV.1972, A. & M. E. Blanchard leg. ( USNM);   Pharr, 8 ♀( AMNH),  Bentsen Rio Grande Valley State Park, 1 Ƌ,  1.XI.1974, B. C. Knudson leg. ( MGCL);   Mercedes, 1 ♀, 1 Ƌ,  30.IV.1959, 1 ♀,  7.V.1959, 1 ♀,  24.VIII.1958, 3 Ƌ, 1 ♀,  31.VIII.1958, 2 ♀,  14.IX.1958, 1 ♀,  21.IX.1958, H. Smalzried leg. ( AMNH);  Cameron, Laguna Atascosa National Wildlife Refuge, 1 Ƌ,  IV.1973, A. & M. E. Blanchard leg., 1 ♀,  25.IV.1995, Don Bowman leg. ( USNM),  Sabal Palm Sanctuary, 1 ♀,  19 –20.VI.1998, E. C. Knudson leg. ( MGCL),   Brownsville, 1 ♀, 1 Ƌ,  7.IV.1919, 1 ♀,  17.IV.1919, 1 ♀, 1 Ƌ,  21.IV.1919, 1 ♀,  23.IV.1919, 1 ♀,  27.IV.1919, 1 ♀,  V.1919, 1 Ƌ,  5.V.1919, 1 ♀,  14.V.1919, 1 Ƌ,  22.V.1919, H. C. Hanson leg. ( USNM).   MEXICO:  Sinaloa: 4 ♀( USNM),  Venadio, 1 Ƌ, USNM 110.114( USNM),  1 ♀, 1919, NHMUK 10921555( NHMUK);   Durango: 3 Ƌ,  VI.1933, Macias leg. ( AMNH);   San Luis Potosí:  Tamazunchale, 1 ♀, 1 Ƌ,  9.IX.1947, P. & C. Vaurie leg., 1 ♀,  20.V.1952, M. Cazier, W. Gertsch& R. Schrammel leg. ( AMNH);   Yucatán:  Cansahcab, 1 Ƌ,  17.VI.1988, R. Segura& Brown leg. ( USNM);   Veracruz:  Jalapa, 1 Ƌ ([ Holo]type of  Meskea horror),  V.1908, R. Muller leg. ( USNM);   Cardel, 1 Ƌ,  4.VI.1940, 2 ♀, 1 Ƌ,  14.VI.1940, J. Camelo G. leg. ( USNM);   Veracruz, 1 Ƌ,  14.III.1986, John D. Gillett leg. ( USNM);   Córdoba, 1 ♀( Paratypeof  Meskea horror) ( USNM);   Orizaba, 1 Ƌ,  I.1945( USNM);   Presidio, 4 ♀,  X.1939, C. C. Hoffmann leg. ( AMNH);   Catemaco, 1 ♀,  29.V.1964, John Pallister leg. ( AMNH);   Colima:  Comala, 2 ♀, 2 Ƌ,  3 –11.VI.2000,  1600m, 19°27’N 103°42’W, V. O. Becker leg., MVOB 121627( MVOB);   Morelos:  Cuernavaca, 2 ♀( Paratypesof  Meskea horror),  VI.1906, 1 ♀( Paratypeof  Meskea horror),  VII.1906( USNM), 1 Ƌ,  VI.1926, Dr. C. Mayer leg. ( ZSM);   Estado de México:  Zacualpan, 2 ♀,  VI.1915, C. C. Hoffmann leg. ( AMNH);   Puebla:  Tehuacán, 1 Ƌ,  27.V.1937, C. C. Hoffmann leg. ( AMNH);   Chiapas:  La Polka, S.of Tonala, 1 ♀,  26.IX.1986, S. J. Weller& M. J. Ryan leg. ( AMNH);   Mapastepec, 2 ♀,  27.V.1933, 6 Ƌ,  29.V.1933, 1 ♀,  30.V.1933, 2 ♀, 2 Ƌ,  31.V.1933, Escalante leg. ( USNM).   GUATEMALA: 1 ♀, B. Brückner S. G. leg. ( MfN);   Retalhuleu:  San Sebastian(L Thiel), 5 ♀( USNM);  Guatemala:   Ciudad de Guatemala, Río Platanos, 1 ♀,  29.X. 1973, 750m, 14°48’N 90°15’W, Erwin& Hevel leg. ( USNM). EL SALVADOR:  San Salvador, 1 Ƌ, 1 ♀,  4.V.1930, Calderon leg. ( USNM), 1 Ƌ, 1951, 1 Ƌ, 2.II.1951, 3 ♀, 5.V.1951, 1 ♀, 16.VII.1951, 1 Ƌ, 3.IX.1951, 1 ♀,   4.IX.1951, 1 ♀, 7.IX.1951, 1 ♀, 1 Ƌ, 9.IX.1951, 1 Ƌ, 17.IX.1951, 1 Ƌ, 25.IX.1951, 1 ♀, 2.X.1951, 1 Ƌ, 5.X.1951, 1 ♀, 7.X.1951, Dr. A. Zilch S. leg., 1 ♀,   18.V.1956, Dr. E. Möhn leg. ( MSFR),  1 ♀,  5.VII.1957, Pab leg. ( AMNH), 1 Ƌ, 1.V.1960, 1 ♀,   15.V. 1960, 600m, B. Bechyné leg. ( ZSM).   COSTA RICA: 1 ♀, 2 Ƌ, P. Biolley leg. ( USNM), 1 Ƌ, Brade leg. ( NHMV); 1 Ƌ, NHMUK 010921502( NHMUK);  Guanacaste: Santa Rosa National Park,  1 ♀,  13 –15.VI.1979, NHMUK 010921616, 1 Ƌ,  20 –22.V.1980, NHMUK 010921531,  1 ♀,  23 –25.V.1980, NHMUK 010921530,  1 ♀, 1 Ƌ,  5 –7.VI.1980, NHMUK 010921550, NHMUK 010921587, 1 Ƌ,  11 –13.VI.1980, NHMUK 010921588,  1 ♀,  22 –27.VI.1981, NHMUK 010921534, D. H. Janzen& W. Hallwachs leg. ( NHMUK); Area de Conservación Guanacaste, 1 ♀, 6.IV. 1989, 160m,  1 ♀,  15.VI. 2010, 295m, D. H. Janzen& W. Hallwachs leg. ( USNM);  Puntarenas: Aranjuez,  1 ♀, IX.  1987, 100m, J. M. Garua leg., NHMUK 010921533( NHMUK);  San José:  1 ♀, 1 Ƌ ( MNHN);  Cartago: Cachi,  2 ♀, NHMUK 010921568, NHMUK 010921569( NHMUK),  1 ♀,  15.VII.1917, NHMUK 10921514( NHMUK).   FIGURE 15. Geographic distribution of  Hepialodesand  Meskeaspecies in North, Central and South America.   PANAMA:  Panamá:  La Cabima,  1 ♀,  16 –30.V.1911, August Busck leg. ( USNM).  VENEZUELA: 1 ♀( MNHN);  Guárico: Hato Masaguaral,  45 kmS Calabozo, 1 ♀, 3 –5.VI.1988, 1 ♀, 8.VI.1988, 1 ♀, 16 –18.IX. 1990, 75m, 8.57N 67.58W, M. Epstein leg., 1 ♀, 20 –21.IX. 1990, 75m, 8.57N 67.58W, M. Epstein &  J. Wilterding leg. ( USNM). PERU:  Lima: San Isidro, 1 ♀,  24.IV.1957,  Salazar leg. ( AMNH). 3032936393 United States of America Bastrop 9 244 1 Texas 3032936346 Cordoba & W. Schaus Mexico Mexico 9 244 1 paratype 3032936422 USNM W. Schaus Mexico Morelos 9 244 1 Morelos paratype 3032936317 1923-07-01 1924-08-30 1923-07-01 USNM C. Poling United States of America Arizona 12 247 7 7 Arizona 3032936403 1923-08-01 1967-06-16 1923-08-01 AMNH C. Poling & J. H. Hessel United States of America 12 247 3 3 Arizona 3032936376 1921-05-05 1923-04-25 1921-05-05 USNM C. Dukes United States of America Mobile 12 247 2 2 Alabama 3032936379 1967-04-30 AMNH R. & B. Taylor United States of America Handsboro Harrison 12 247 1 1 Mississippi 3032936396 [628,1432,1940,1967] 1991-05-10 USNM H. Xeroobien United States of America Diamondhead Hanckock 12 247 1 1 Mississippi 3032936362 [151,1176,1976,2003] 2017-04-28 MGCL J. Hayden United States of America Rock Hill Prsv. Chipley 12 247 1 1 Florida 3032936363 1968-05-16 1987-05-24 1968-05-16 MGCL H. V. Weems Jr. & H. Greenbaum & H. O. Baggett & Charles Stevens United States of America 13 248 Torreya State Park Liberty 12 247 4 4 Washington 3032936432 [771,1432,150,177] NHMUK United States of America Tallahassee Leon 13 248 NHMUK 10921563 1 1 Washington 3032936427 [151,896,187,213] 1941-05-13 MGCL Shirley Hills United States of America Pensacola Escambia 13 248 1 1 Washington 3032936347 [907,1432,187,213] AMNH United States of America Baia de Biscayne Miami-Dade 13 248 1 1 Washington 3032936377 [151,907,223,249] 1910-03 1927-02 1910-03 USNM United States of America Paradise Key 13 248 3 3 Washington 3032936447 1924-04-13 1924-04-29 1924-04-13 MfN L. J. Bottimer & H. C. Hanson & Friedrich United States of America Texas 13 248 14 14 Texas 3032936312 [267,1287,295,321] 1964-05-08 USNM A. & M. E. Blanchard United States of America Lake Brownwood State Park Brown 13 248 1 1 Texas 3032936442 1988-05-08 1990-05-02 1988-05-08 MGCL E. C. Knudson & Linwood C. Dow United States of America San Jacinto San Houston National Forest Coldspring 13 248 2 2 Texas 3032936328 [374,1175,366,393] 1966-05-20 USNM A. & M. E. Blanchard United States of America Junction Kimble 13 248 4 4 Texas 3032936319 USNM United States of America Shovel Mountain Burnet 13 248 1 1 Texas 3032936368 [373,1221,402,429] 1969-05-11 USNM A & M. E. Blanchard United States of America Conroe Montgomery 13 248 1 1 Texas 3032936324 1919-05-04 1919-05-10 1919-05-04 AMNH Holo United States of America Bastrop Harris Bastrop 13 248 1 1 Texas 3032936358 1919-04-14 1919-05-10 1919-04-14 USNM H. C. Hanson United States of America 13 248 5 5 Texas 3032936434 [647,1205,474,501] 1919-05-12 AMNH United States of America Wallisville Chambers 13 248 1 Texas 3032936407 1919-05-09 1919-05-19 1919-05-09 NMHN C. P. Heinrich & A. Busck United States of America 13 248 2 2 Texas 3032936425 [268,1359,546,573] 1923-04-11 1923-04-25 1923-04-11 AMNH United States of America Lake Charlotte 13 248 2 2 Texas 3032936401 1923-04-17 1923-04-26 1923-04-17 NMHN L. J. Bottimer United States of America 13 248 14 14 Texas 3032936446 1992-05-01 2005-05-07 1992-05-01 MGCL C. Bordelon & E. C. Knudson & Bordelon & Knudson United States of America ConCan Uvalde 13 248 4 4 Texas 3032936366 1992-04-30 1994-05-09 1992-04-30 USNM Ron Leuschner & Kurt Leuschner United States of America Rio Frio 13 248 1 1 Texas 3032936309 1928-04-28 USNM H. B. Parks United States of America Bexar 13 248 1 Texas 3032936331 1976-05-16 1977-05-17 1976-05-16 MGCL E. C. Knudson & William W. & Nadine M. United States of America San Antonio 13 248 1 1 Texas 3032936373 [354,1432,763,789] 1977-04-01 MGCL William W. & Nadine M. United States of America Bexar Loop Camp Bullis Road 13 248 1 1 Texas 3032936398 [151,588,799,825] EHRS United States of America Ebony Hill Res Station Bexar 13 248 1 Texas 3032936415 [599,1315,798,825] 1985-04-25 MGCL Roy O. Kendall & C. A. Kendall United States of America 13 248 1 Texas 3032936436 1963-04-20 1965-05-19 1963-04-20 USNM A. & M. E. Blanchard United States of America San Patricio Welder National Wildlife Refuge Sinton 13 248 6 6 Texas 3032936337 1972-04-07 USNM A. & M. E. Blanchard United States of America Santa Ana National Wildlife Refuge Hidalgo 13 248 1 1 Texas 3032936394 [475,722,907,933] AMNH United States of America Pharr 13 248 8 8 Texas 3032936381 1974-11-01 MGCL B. C. Knudson United States of America Bentsen Rio Grande Valley State Park 13 248 1 Texas 3032936437 1958-08-24 1959-05-07 1958-08-24 AMNH H. Smalzried United States of America Mercedes 13 248 7 7 Texas 3032936409 1973-04 1995-04-25 1973-04 USNM A. & M. E. Blanchard & Don Bowman United States of America Laguna Atascosa National Wildlife Refuge Cameron 13 248 1 1 Texas 3032936416 1998-06-19 1998-06-20 1998-06-19 MGCL E. C. Knudson United States of America Sabal Palm Sanctuary 13 248 1 1 Texas 3032936448 1919-04-07 1919-05-22 1919-04-07 USNM H. C. Hanson United States of America Brownsville 13 248 7 7 Texas 3032936348 [199,587,1159,1185] USNM Mexico Sinaloa 13 248 4 4 Sinaloa 3032936306 [600,1061,1159,1185] USNM Mexico Venadio 13 248 USNM 110.114 1 3032936426 NHMUK Mexico 13 248 NHMUK 10921555 1 1 3032936406 [291,813,1195,1221] 1933-06 AMNH Macias Mexico Durango 13 248 1 Durango 3032936310 1947-09-09 1952-05-20 1947-09-09 AMNH P. & C. Vaurie & M. Cazier & W. Gertsch & R. Schrammel Mexico Tamazunchale 13 248 2 2 San Luis Potosi 3032936384 1988-06-17 USNM R. Segura & Brown Mexico Cansahcab 13 248 1 Yucatan 3032936419 1908-05 USNM R. Muller Mexico Holo Jalapa 13 248 1 Veracruz de Ignacio de la Llave 3032936420 [434,1286,1302,1329] 1940-06-04 1940-06-14 1940-06-04 USNM Camelo G. Mexico Cardel 13 248 2 2 3032936311 1986-03-14 USNM John D. Gillett Mexico Veracruz 13 248 1 Veracruz de Ignacio de la Llave 3032936321 [663,1263,1338,1365] USNM Mexico Cordoba 13 248 1 1 paratype 3032936441 1945-01 USNM Mexico Orizaba 13 248 1 paratype 3032936349 [345,954,1374,1401] 1939-10 AMNH C. C. Hoffmann Mexico Presidio 13 248 4 4 paratype 3032936305 1964-05-29 AMNH John Pallister Mexico Catemaco 13 248 1 1 paratype 3032936326 2000-06-03 2000-06-11 2000-06-03 MVOB V. O. Becker Mexico 1600 19.45 Comala 1271 -103.7 13 248 MVOB 121627 2 2 Colima paratype 3032936360 1906-06 1906-07 1906-06 USNM Mexico Cuernavaca 13 248 3 3 Morelos paratype 3032936353 [470,913,1483,1509] 1926-06 ZSM Dr. C. Mayer Mexico 13 248 1 paratype 3032936333 1915-06 AMNH C. C. Hoffmann Mexico Zacualpan 13 248 2 2 Mexico paratype 3032936336 [514,1281,1518,1545] 1937-05-27 AMNH C. C. Hoffmann Mexico Tehuacan 13 248 1 Puebla paratype 3032936304 1986-09-26 AMNH La Polka, S. & S. J. Weller & M. J. Ryan Mexico Tonala 13 248 1 1 Chiapas paratype 3032936390 1933-05-27 1933-05-31 1933-05-27 USNM Escalante Mexico Mapastepec 13 248 5 5 paratype 3032936375 [199,778,1627,1654] MfN Bruckner S. G. GUATEMALA 13 248 1 1 Guatemala 3032936372 [789,1367,1626,1654] USNM Thiel San Sebastian 13 248 5 5 Retalhuleu 3032936380 1973-10-29 USNM Erwin & Hevel 14.8 Rio Platanos 1286 -90.25 13 248 1 1 Guatemala 3032936352 [265,1131,1698,1725] 1930-05-04 USNM Calderon San Salvador 13 248 1 1 Guatemala 3032936314 [799,974,1734,1761] 1951-09-04 USNM Calderon true San Salvador 13 248 1 1 Guatemala 3032936378 1956-05-18 MSFR Dr. E. Mohn 13 248 1 Guatemala 3032936359 [457,857,1806,1833] 1957-07-05 AMNH Pab 13 248 1 1 Guatemala 3032936391 1960-05-15 ZSM B. Bechyne 13 248 1 Guatemala 3032936382 USNM, NHMV, NHMUK D, P. Biolley & Brade Costa Rica Santa Rosa National Park 13 248 NHMUK 010921502 1 1 Guanacaste 3032936392 [733,1233,1914,1941] 1979-06-13 1979-06-15 1979-06-13 NHMUK Costa Rica 13 248 NHMUK 010921616, 1 1 1 3032936344 1980-05-20 1980-05-22 1980-05-20 NHMUK Costa Rica 13 248 NHMUK 010921531 1 3032936410 [410,891,1950,1977] 1980-05-23 1980-05-25 1980-05-23 NHMUK Costa Rica 13 248 NHMUK 010921530 1 1 3032936383 1980-06-05 1980-06-07 1980-06-05 NHMUK Costa Rica 13 248 NHMUK 010921550, NHMUK 010921587, 1 1 1 3032936421 [422,878,1986,2013] 1980-06-11 1980-06-13 1980-06-11 NHMUK Costa Rica 13 248 NHMUK 010921588 1 3032936318 1981-06-22 1981-06-27 1981-06-22 NHMUK D. H. Janzen & W. Hallwachs Costa Rica 14 249 Guanacaste 13 248 NHMUK 010921534 1 1 Guanacaste 3032936430 2010-06-15 USNM D. H. Janzen & W. Hallwachs Costa Rica Aranjuez 14 249 1 1 Puntarenas 3032936330 NHMUK J. M. Garua Costa Rica 1987 San Jose 14 249 NHMUK 010921533 1 1 San Jose 3032936323 [712,1119,223,249] MNHN Costa Rica Cachi 14 249 1 1 Cartago 3032936334 NHMUK Costa Rica 14 249 NHMUK 010921568, NHMUK 010921569 2 2 3032936408 [535,1102,258,285] 1917-07-15 NHMUK Costa Rica 14 249 NHMUK 10921514 1 1 3032936332 [199,576,150,177] La Cabima Panama Panama 15 250 1 Panama 3032936355 [589,1130,151,177] 1911-05-16 1911-05-30 1911-05-16 USNM August Busck Panama 15 250 1 1 3032936343 [199,1050,186,213] MNHN Venezuela Hato Masaguaral 15 250 1 1 Guarico 3032936302 1957-04-24 USNM J. Wilterding Peru San Isidro 15 250 1 1 Lima 3032936445 [680,926,295,321] AMNH Salazar Peru 15 250 1