Croton squamiger Croton squamiger var. obtusifolius Croton squamiger Oxydectes jennyana Croton jennyanus Oxydectes squamigera Croton squamiger A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands Berry, Paul E. Kainulainen, Kent Ee, Benjamin W. van PhytoKeys 2017 2017-11-15 90 1 87 47987780-49AC-5828-893B-17E8D3E86C7A Gris ex Baill., Adansonia 1: 160. 1861, as ' jennyanum' Gris ex Baill., Adansonia 1: 160. 1861, as ‘ jennyanum’ Magnoliopsida Euphorbiaceae Croton CoL Plantae Croton jennyanus Malpighiales 40 41 Tracheophyta species jennyanus   Croton squamigerBaill., Adansonia 1: 168. 1861, as ' squamigerum'. Type. Madagascar. Prov. Antsiranana: sin. loc., s.d., J.M.C. Richard 576(lectotype, designated here: P [P00133578]!; isolectotype: P [P00133579]!). Madagascar. Prov. Antsiranana: Diego-Suareset baie de Rigny, 1837, J.M.C. Richard 176(syntype: P [P00133577]!). Madagascar. Prov. Antsiranana: Cap d'Ambre, 1847-1852, L.H. Boivin 2658(syntype: P [P00133273]!).  Croton squamiger var. obtusifolius Muell.Arg. in A.P.de Candolle, Prodr. 15(2): 523. 1866, nom. inval. Type. Based on Croton squamigerBaill.  Oxydectes jennyana(Gris ex Baill.) Kuntze, Revis. Gen. Pl. 2: 612. 1891. Type. Based on Croton jennyanusGris ex Baill.  Oxydectes squamigera(Baill.) Kuntze, Revis. Gen. Pl. 2: 613. 1891. Type. Based on Croton squamigerBaill.   Type.   Madagascar. Prov. Antsiranana[Diana Region]: vers les bas-fonds humides, lorsqu'on s'elevepar les torrents au dessus des montagnes qui se trouvent au nord du port Lugwata[Diego Suarez], 1833,  J.P. Goudots.n.( holotype: G [G00434419]!; isotypes: P [P00133278]!, P [P00154519]!).  Habit and distribution. Shrubs; northern and western Madagascar (Antsiranana, Mahajanga).  Notes. We follow here the precedent of Leandri (1939)and Radcliffe-Smith (2016)in treating  Croton squamigeras a synonym of  C. jennyanus, but we differ in treating  C. hypochalibaeusas a distinct, more highland species rather than as another synonymof  C. jennyanus(see Kainulainen et al., 2016). According to our interpretation,  C. jennyanusis restricted to lower elevations in northern Madagascar (Montagne des Francais, Sahafary, Daraina, Ankarana), as well as in far midwestern Madagascar on or near tsingy formations (Bemaraha). Concerning the type of  Croton squamiger, three other sheets of Boivin 2658correspond to  C. brevispicatus, so this is clearly a mixed collection. In the description of  C. squamiger, Baillon (1861)divided the species into two infraspecific taxa, a and b. These were subsequently named by  Mueller(1866)as C. squamigerus var. obtusifolius(the typical variety) and C. squamigerus var. acutifolius, respectively. The latter has been recognized here to be synonymous with  C. hypochalibaeus. We have corrected the termination of the specific epithet to "  Croton squamiger" [masc. gender, Adj. group A nom., see Stearn (1992: 91) and Art. 23.5 and Art. 60.9 Ex. 24 ( ICN 2012;  Croton ciliato-glanduliferumOrtega corrected to  C. ciliatoglandulifer).  Radcliffe-Smith (2016)listed  Croton jennyanusas being present in the Comoros Archipelago. The three collections he cited [ Pascal 928(K, P) from Mayotte, and Floret 1241(P) and 1249(P) from Moheli] all correspond to  C. humblotii. 2017-01-01 2017-12-31 2017-01-01 J. P. Goudot Madagascar port Lugwata 1 Antsiranana holotype