Xantholinus insulatus Cameron 1922: 115 Xantholinus solitarius Blackwelder 1943: 486 TRIBESTAPHYLININI Belonuchus amplus Blackwelder 1943: 425 Belonuchus antiguae Blackwelder 1943: 434 Belonuchus bugnioni Fauvel 1901: 86 Belonuchus eximius Bernhauer 1917b: 105 Belonuchus dominicus Blackwelder 1943: 431 Belonuchus gagates Erichson 1840: 424 America Belonuchus mundus Erichson 1840: 425 Cafius bistriatus (Erichson) 1840: 502 Philonthus Mexico Neobisnius funerulus Cameron 1922: 119 Neobisnius humilis (Erichson) 1840: 512 Philonthus Philonthus trepidus Erichson 1840: 489 Philonthus varians (Paykull) 1789: 45 Staphylinus America Philonthus waterhousei Cameron 1922: 120 SUBTRIBEHYPTIOMINA Holisusguildingii Erichson 1839: 300 The beetles of the Lesser Antilles (Insecta, Coleoptera): diversity and distributions Peck, Stewart B. Insecta Mundi 2016 2016-02-12 2016 460 1 360 8VR6D Latreille 1802 [192,614,291,315] Insecta Staphylinidae Animalia Coleoptera 83 78 Arthropoda subFamily Staphylininae    Diochus nanusErichson 1839: 301; Blackwelder 1943: 455; Herman 2001: 2446; Ivieet al. 2008b: 243; Franket al. 2011: 8. =  Diochus perplexusCameron 1922: 116of Grenada. =  Diochus apicipennisCameron 1922: 116of St. Vincent. =  Diochus antennalisCameron 1922: 117of Grenada. Distribution. Barbados, Caymans, Cuba, Grenada, Guadeloupe, Hispaniola, Jamaica, Montserrat, Mustique, Puerto Rico, St. Croix, St. Lucia, St. Vincent. Mexicoto Colombia, Venezuelaand Trinidadto Braziland Argentina; widespread Antillesand Latin America.  TRIBEXANTHOLININI   Lithocharodes claviscapa(Cameron) 1922: 114(  Somoleptus); Blackwelder 1943: 497;  Herman2001: 3694.  Distribution. Grenada( typelocality), St. Vincent; Lesser Antillesendemic.   Microlinus pusio(LeConte) 1880: 171(  Leptolinus); Ivie et al. 2008b: 243. Distribution. Montserrat. Se USA(SC, FL); widespread Antillesand North and/or Central America.   Neohypnus attenuatus(Erichson) 1839: 330(  Xantholinus); Blackwelder 1943: 478; Herman 2001: 3708; Ivieet al. 2008b: 244. Distribution. Antigua, Barbados, Cuba, Dominica, Grenada, Guadeloupe, Jamaica, Montserrat, PuertoRico, St. Kitts, St. Thomas, St. Vincent. USA(CA-TX-FL), Mexico, Venezuela, Trinidad, Tobago, Brazil, Paraguay, Argentina; St. Helena(introduced); widespread New World.   Neohypnus humeralis(Erichson) 1839: 327(  Xantholinus); Blackwelder 1943: 479; Herman 2001: 3799; Ivieet al. 2008b: 244. Distribution. Antigua, Cuba, Dominica, Hispaniola, Montserrat, Puerto Rico, St. Croix, St John, St. Vincent; widespread Antillesendemic.   Neohypnus illucens(Erichson) 1839: 315(  Xantholinus); Blackwelder 1943: 488; Herman 2001: 3800; Ivie et al. 2008b: 244. Distribution. Dominica, Grenada, Guadeloupe, Montserrat, Puerto Rico, St. Lucia, St. Thomas, St. Vincent. Colombia, Venezuela, Tobago, Trinidad; widespread Antillesand Latin America.   Neoxantholinus filarius(Erichson) 1839: 334(  Leptacinus); Blackwelder 1943: 506 (  Oligolinus); Herman 2001: 3716. Distribution. St. Vincent. Mexicoto Colombia, Venezuelato Ecuador, Brazil; the Lesser Antillesand Latin America.   Neoxantholinus hubbardi(Blackwelder) 1943: 505(  Oligolinus); Herman 2001: 3717; Ivieet al. 2008b: 244. Distribution. Montserrat; singleisland endemic.   Phacophallus parumpunctatus(Gyllenhal) 1827: 481(  Staphylinus); Blackwelder 1943: 494 (  Leptacinus); Tucker 1952: 341; Bennettand Alam 1985: 21; Herman 2001: 3738. =  Phacophallus fauveliCameron 1922: 114of Grenadaandof St. Vincent. Distribution. Antigua, Bahamas, Barbados, Cuba, Grenada, Jamaica, Puerto Rico, St. Kitts, St. Vincent. Widespread: USAand Canada; Africa; Asia; Pacific Islands; cosmopolitan; native to Palearctic region; introduced to New World, introduced to the Lesser Antilles; now widely distributed in North America, but seemingly not in Central and South America (Smetana 1982: 108). Notes.In decaying organic matter, especially in synanthropic situations; in chicken manure and cow dung.   Thyreocephalus subtilisSharp 1885a: 499; Blackwelder 1943: 491; Herman 2001: 3767. Distribution. St. Vincent. Panama, Colombia, Venezuela, Peru, Brazil; the Lesser Antillesand Latin America.   Xantholinus brunneipennisErichson 1839: 327; new species record. Distribution. Grenada(based on 1 specimenin FMNHdetermined by Bernhauer). Colombia; the Lesser Antilles and Latin America.    Xantholinus insulatusCameron 1922: 115; Blackwelder 1943: 476;  Herman2001: 3801.  Distribution. Grenada( typelocality), Mustique; Grenadapaleo-island endemic.   Xantholinus solitariusBlackwelder 1943: 486; Herman 2001: 3823. Distribution. Guadeloupe; single island endemic.  TRIBESTAPHYLININI   SUBTRIBETANYGNATHININA   Atanygnathus heterocerusCameron 1922: 123; Blackwelder 1943: 471: Herman 2001: 3561. Distribution. Grenada, St. Vincent; Lesser Antilles endemic. Notes. Ivie et al. (2008b: 243) list two unidentified species in this genus from Montserrat.   Atanygnathus laticollis(Erichson) 1839: 289(  Tanygnathus);  Blackwelder1943: 472: Herman2001: 3562. Distribution. Antigua, Cuba, Hispaniola, Puerto Rico, St. Vincent ( typelocality). Trinidad; widespread Antillesand Latin America.  SUBTRIBEXANTHOPYGINA   Xanthopygus pexus(Motschulsky) 1858: 667(  Belonuchus); Blackwelder 1943: 450; Herman 2001: 3608. Distribution. St. Vincent. Trinidad, Colombiaand Venezuela, Guyana, Brazil, Paraguay; the Lesser Antillesand Latin America.   Xenopygus analis(Erichson) 1840: 495(  Philonthus); Blackwelder 1943: 453 (  Philothalpus); Irmler 1979; 32; Herman 2001: 3610. Distribution. Grenada(implied on map by Irmler 1979: 32, not mentioned in text). Mexicoto Colombia, Trinidad, to Bolivia, Brazil; widespread Antilles and Latin America.  SUBTRIBEPHILONTHINA    Belonuchus amplusBlackwelder 1943: 425; Herman 2001: 2519. Distribution. St. Lucia; single island endemic. Note.Ivie et al. (2008b: 243) reportthree unidentified species in thisgenus from Montserrat.   Belonuchus antiguaeBlackwelder 1943: 434; Herman 2001: 2519. Distribution. Antigua; singleisland endemic.   Belonuchus bugnioniFauvel 1901: 86;  Blackwelder1943: 428: Herman2001: 2521. =  Belonuchus eximiusBernhauer 1917b: 105of Guadeloupe. Distribution. Guadeloupe, Martinique( typelocality); Lesser Antillesendemic.    Belonuchus coelestinusBernhauer 1908: 330; Blackwelder 1943: 426; Herman 2001: 2521. =  Belonuchus coeruleusCameron 1922: 122of St. Vincent. =  Belonuchus smithiCameron 1922: 122, manuscript name of Fauvel. Distribution. Grenada, St. Vincent( typelocality); Lesser Antilles endemic.   Belonuchus cognatusSharp 1885a: 429; Blackwelder 1943: 427; Herman 2001: 2522. Distribution. St. Vincent. Belizeto Venezuela, Trinidad, Argentina; the Lesser Antilles and Latin America. Plate3.    Belonuchus dominicusBlackwelder 1943: 431; Herman 2001: 2524. Distribution. Dominica; single island endemic.   Belonuchus gagatesErichson 1840: 424; Ivie et al. 2008b: 243; Turnbow and Thomas 2008: 52; Frank et al. 2011: 8. Distribution. Bahamas, Caymans, Cuba, Jamaica, Hispaniola, Montserrat, Puerto Rico, St. John, St. Thomas. USA(FL); widespread Antillesand North and/or Central America.   Belonuchus mundusErichson 1840: 425; Blackwelder 1943: 430; Herman 2001: 2530. Distribution. Grenada, St. Lucia, St. Vincent; Lesser Antillesendemic.   Cafius bistriatus(Erichson) 1840: 502(  Philonthus); Blackwelder 1943: 43; Ramos 1946: 33; Franket al. 1986: 149, 158; Woodruff et al. 1998: 42; Herman 2001: 2569, Ivie et al. 2008b: 243; Turnbow and Thomas 2008: 52. Distribution. Antigua, Bahamas, Barbados, Carriacou, Cuba, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Mona, Montserrat, PuertoRico, St. Croix, St. John, St. Kitts, St. Lucia, St. Thomas. Canada(NB, NS, PQ), USA(ME-FL-TX), Mexicoto Trinidad, Tobago; widespread New World. Note.Found under seaweed and drift on beaches.    Cafius caribeanusBierig 1934a: 68; Blackwelder 1943: 437; Frank et al. 1986: 149; Woodruff et al. 1998: 42; Herman 2001: 2571. Distribution. Antigua, Carriacou, Cuba, Dominica, Grenada, Guadeloupe, Jamaica, PuertoRico, St. Croix. USA(FL), Mexico, Panama, Venezuela, Trinidad, Brazil; widespread New World. Note.Found under seaweed and drift on beaches.   Cafius subtilisCameron 1922: 121; Blackwelder 1943: 436; Ramos 1946: 33; Frank et al. 1986: 149; Herman 2001: 2578; Ivieet al. 2008b: 243. Distribution. Antigua, Cuba, Dominica, Guadeloupe, Jamaica, Mona, Montserrat, Puerto Rico, St. Croix, St. Kitts. USA(FL); widespread Antillesand North and/or Central America. Note.Found under seaweed and drift on beaches.   Gabronthus thermarum(Aubé) 1850: 316(  Philonthus); Blackwelder 1943: 403; Frank 1983: 476; Smetana 1995: 481; Herman 2001: 2670; Perez-Gelabert 2008: 91. Distribution. Antigua, Grenada, Guadeloupe, Hispaniola, Jamaica, Puerto Rico, St. Croix, St. Kitts. USA(e), Mexico, Costa Rica, French Guiana, Africa, Europe, Asia; Indian and Pacific ocean islands; widespread New World. Plate5.    Neobisnius funerulusCameron 1922: 119; Blackwelder 1943: 1131; Frank 1981: 34;  Herman2001: 2705.  Distribution. St. Lucia, St. Vincent( typelocality); the Lesser Antillesendemic. [  Neobisnius humilis(Erichson) 1840: 512(  Philonthus); Franket al. 2011: 8. Distribution. Caymans, Cuba, Hispaniola, Jamaica, Puerto Rico, Virgin Islands; not Grenada; earlier the Lesser Antilles records in Blackwelder 1943 are synonyms and were removed by Frank 1981.]    Neobisnius ludicrus(Erichson) 1840: 514(  Philonthus); Frank 1981: 20; Herman 2001: 2708; Distribution. Barbados, Grenada, St. Lucia, St. Vincent. USA(SC-FL-TX), Mexicoto Colombia, Venezuela, Trinidadto Brazil; widespread New World. Plate7.   Paederomimus cribricollis(Erichson) 1840: 487(  Philonthus); Blackwelder 1943: 396; Herman 2001: 2725. Distribution. Grenada. Mexico?, Colombia, Venezuela, Trinidad, Peru; the Lesser Antilles and Latin America.   Paederomimus interjectusBernhauer 1908: 333; Blackwelder 1943: 397; Herman 2001: 2726. =  Paederomimus smithiCameron 1922: 121of St. Vincent. Distribution. St. Vincent; singleisland endemic.   Philonthus discoideus(Gravenhorst) 1802: 38(  Staphylinus); Blackwelder 1943: 407; Smetana 1995: 184; Herman 2001: 2807; Ivieet al. 2008b: 244. Distribution. Antigua, Cuba, Grenada, Guadeloupe, Hispaniola, Jamaica, Montserrat, Puerto Rico, St. Croix, St. Kitts. North America(widespread), Mexicoto Venezuela; Atlantic Islands; Europe; Asia; Africa; widespread New World.   Philonthus figulusErichson 1840: 464; Blackwelder 1943: 409; Herman 2001: 2822. Distribution. Cuba, Puerto Rico, St. Vincent. Mexicoto Colombia, Trinidad, Venezuelato Brazil, Argentina; widespread Antillesand Latin America.   Philonthus flavolimbatusErichson 1840: 471; Blackwelder 1943: 414; Smetana 1995: 200; Herman 2001; 2824. Distribution. Carriacou, Grenada, Martinique. Canada, USA(widespread), Mexicoto Venezuelato Argentina; widespread New World.   Philonthus havaniensis(Laporte) 1835: 116(  Staphylinus); Blackwelder 1943: 418; Ramos 1946: 33; Herman 2001: 2836; Ivieet al. 2008b: 244; Franket al. 2011: 8. Distribution. Antigua, Cuba, Caymans, Hispaniola, Jamaica, Mona, Montserrat, PuertoRico, St. Croix, St. John; widespread Antillesendemic.   Philonthus hepaticusErichson 1840: 451; Blackwelder 1943: 401; Bennett and Alam 1985: 21; Smetana 1995: 175; Herman 2001: 2837; Ivieet al. 2008b: 244; Turnbowand Thomas 2008: 53. =  Philonthus vilisErichson 1825: 610; Fleutiauxand Sallé 1890: 379 of Guadeloupe. Distribution. Antigua, Bahamas, Barbados, Cuba, Dominica, Grenadines, Guadeloupe, Hispaniola, Jamaica, Les Saintes, Montserrat, PuertoRico, St. Croix, St. Kitts, St. Thomas, St. Vincent. Canada(NB), USA(widespread), Mexicoto Panama, Colombia, Tobago, Trinidad, Venezuela, Argentina, Chile; widespread New World; Australia, New Zealand.   Philonthus longicornisStephens 1832: 237; Blackwelder 1943: 410; Smetana 1995: 245; Herman 2001: 2866; Ivieet al. 2008b: 244. Distribution. Antigua, Cuba, Grenada, Hispaniola, Montserrat, Puerto Rico, St. Kitts. Tropicopolitan; widespread New World.    Philonthus trepidusErichson 1840: 489; Smetana 1994: 345. Distribution. Antigua, Puerto Rico, St. Croix, St. John; widespread Antillesendemic.   Philonthus varians(Paykull) 1789: 45(  Staphylinus); Blackwelder 1943: 412; Smetana 1995: 235; Herman 2001: 2989. Distribution. Cuba, St. John, St. Vincent. Widespread; Europe, Asia, Africa, North America; widespread New World.    Philonthus ventralis(Gravenhorst) 1802: 174(  Staphylinus); Fleutiaux and Sallé 1890: 380; Blackwelder 1943: 404; Ramos 1946: 33; Woodruff et al. 1998: 43; Bennettand Alam 1985: 21; Smetana 1995: 190; Herman 2001: 2996; Ivie et al. 2008b: 244; Frank et al. 2011: 9. Distribution. Antigua, Barbados, Caymans, Cuba, Culebra, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Mona, Montserrat, PuertoRico, St. Croix, St. Kitts, St. Lucia, St. Thomas, St. Vincent. USA(widespread), Mexico, Costa Rica, French Guiana, Tobago, Trinidad; Africa; Asia; Europe; cosmopolitan; widespread New World.    Philonthus waterhouseiCameron 1922: 120; Blackwelder 1943: 417;  Herman2001: 3003.  Distribution. Grenada, St. Vincent ( typelocality); Lesser Antillesendemic.  SUBTRIBEHYPTIOMINA    Holisus debilisErichson 1839: 300; Blackwelder 1943: 401; Herman 2001: 2512; Turnbow and Thomas 2008: 52. Distribution. Bahamas, Cuba, Hispaniola, PuertoRico, St. John, St. Lucia, St. Vincent; widespread Antilles endemic. Note.Ivie et al. (2008b: 243) report two unidentified species in this genus from Montserrat.    HolisusguildingiiErichson 1839: 300; Blackwelder 1943: 462: Herman 2001: 2512. Distribution. Antigua, Puerto Rico, St. Lucia, St. Vincent ( typelocality); widespread Antillesendemic. Grenada Herman Distribution. 83 78 1 FMNH Grenada Bernhauer Grenada 83 78 1 Grenada Herman Distribution. 83 78 1 Antigua and Barbuda Hispaniola Blackwelder Herman 84 79 1 Guadeloupe Blackwelder Herman 84 79 1 St. Lucia, St. Vincent Herman Distribution. 85 80 1 Grenada Herman Distribution. 85 80 1 Antigua, Puerto Rico, St. Lucia Antigua, Puerto Rico, St. Lucia 86 81 1