Rhyacobates recurvus Andersen & Chen, 1995: 59 Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae) Leng, Zhaoqi Tran, Anh Duc Ye, Zhen European Journal of Taxonomy 2023 2023-09-28 893 1 1 73 4SNHT Andersen & Chen, 1995 Andersen & Chen 1995 [515,1073,264,292] Insecta Gerridae Rhyacobates Animalia Hemiptera 53 54 Arthropoda species recurvus   Figs 3N, 4K, 5M, 7D, 8J, 14, 31    Rhyacobates recurvus Andersen & Chen, 1995: 59, figs 16–17 (original description).    Diagnosis Body length of apterous females 7.06–8.71, of apterous males 5.42–5.91. Female: pronotum mainly black with a median brownish-yellow spot; mesonotum and metanotum mainly black with a median brownish-yellow stripe ( Figs 3N, 31A); abdomen relatively short, with posterior part distinctly curved dorsad to oblique or nearly vertical position ( Figs 8J, 31C); abdominal mediosternite VII tapering caudad, posterior margin with a median process; apex of the process pointed and hook-shaped ( Fig. 31D, F). Male: middle trochanter with one spine; middle femur with scattered small spines, not arranged in distinct row ( Fig. 31I); length of middle tibia ca 1.6 times length of hind tibia; proctiger laterally with rounded lobes ( Figs 7D, 31J); paramere relatively slender, strongly curved at basal fourth, middle part thickened, distal part tapering towards narrowly rounded apex ( Figs 8J, 31J).    Material examined  Non-type specimens   CHINA–  Hubei Province• 7 ♁♁, 5 ♀♀(apterous); Huang-gang City, Ying-shan County, Jiu-gong Mountain; 29°3′25.6″ N, 107°5′55.1″ E;  130 ma.s.l.;  11 Jul. 2022; Zhao-qi Lengand Zhen Yeleg.; NKUM. –   Jiangxi Province• 1 ♁, 2 ♀♀(apterous); Nan-chang City;  2 Jun. 1919; IZAS. –   Zhejiang Province• 3 ♀♀(apterous); Jin-hua City, Pan-an County, Da-pan Mountain; 28°58′30.1″ N, 120°31′36.4″ E;  29 Jul. 2015; Wen-bo Yileg.; NKUM.    Supplemental description  Apterous female  MEASUREMENTS. Body length 7.06–8.71, width 2.63–2.90, head width 1.42, interocular width 0.53, eye length (dorsal view) 0.60; relative lengths of antennal segments I–IV: 3.95: 1.05:0.93:0.78; pronotum: length 0.74, width 1.52;mesonotum: length2.44, width 2.63; metanotum:length 0.69, width 2.34; abdomen length (ventral view) 1.96; abdominal sternum VII: length 0.90, width 1.12; abdominal mediotergite I: length 0.28, width 1.09; relative lengths of leg segments (femur:tibia: tarsal segment I:tarsal segment II): fore leg: 4.10: 3.65:1.67:0.83, middle leg: 12.02:6.95: 2.91: 0.38, hind leg: 12.35:5.05:0.16:0.18.  COLORATION. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum and metanotum mainly black with a median brownish-yellow stripe ( Figs 3N, 31A). Abdominal tergum I with yellowish spots on each side or completely yellow. Connexivum dorsally brownish-yellow. Mesosternum chiefly blackish with a median subtriangular yellowish spot ( Fig. 31B). Abdominal venter light yellow.  ABDOMEN. Abdomen relatively short, with posterior part distinctly curved dorsad to oblique or nearly vertical position ( Figs 5M, 31C). Connexiva erect on abdominal segments I–II, converging along dorsal midline of segments III–VI, reflexed above terminal terga ( Fig. 31A). Abdominal segment VII elongate, ventral margin longer than dorsal margin in lateral view ( Figs 5M, 31C), nearly as long as three preceding abdominal segments together, not completely enclosing genital segments ( Fig. 31D); laterosternites of connexivum VII expanded dorsad, folded mesad ( Fig. 31A), meeting in middle of abdominal dorsum or overlapping each other ( Fig. 31E). Abdominal sternum VII tapering caudad, posterior margin with a median process; apex of the process pointed and hook-shaped ( Fig. 31D, F).   Fig. 31.Morphological features of  Rhyacobates recurvus Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm.  Apterous male  MEASUREMENTS. Body length 5.42–5.91, width 1.73–2.01, head width 1.23, interocular width 0.41, eye length (dorsal view) 0.49; relative lengths of antennal segments I–IV: 2.93: 0.93: 0.97: 0.73; pronotum: length 0.64, width 1.27; mesonotum: length 1.78, width 1.83; metanotum: length 0.63, width 1.73; abdomen length (ventral view) 1.91; abdominal sternum VII: length 0.42, width 0.79; abdominal mediotergite I: length 0.21, width 0.56; relative lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 3.08: 2.64: 0.88: 0.54, middle leg: 9.64: 4.87: 2.17: 0.32, hind leg: 10.06: 3.10: 0.13: 0.16.  COLORATION. Median black spot of head posteriorly bifurcate. Mesonotum mainly black with a median brownish-yellow stripe. Metanotum chiefly blackish, without yellow markings ( Figs 4K, 31G). Connexivum dorsally blackish. Mesosternum chiefly blackish with a median subtriangular yellowish spot, narrower than that of female ( Fig. 31H). Abdominal venter light yellow.  LEG. Middle trochanter with one spine; middle femur with scattered small spines, not arranged in distinct row ( Fig. 31I).  GENITALIA. Abdominal segment VIII ventro-laterally impressed. Pygophore large, ovate. Proctiger with rounded lobes laterally ( Figs 7D, 31J). Paramere relatively slender, strongly curved at basal fourth, middle part thickened, distal part tapering towards narrowly rounded apex ( Figs 8J, 31J).    Distribution  China: Hubei, Zhejiang(first records); Jiangxi( Andersen & Chen 1995).    Comparative notes   Rhyacobates recurvusis most similar to  R. turgidus sp. nov.; see comparative notes under  R. turgidus.   Remarks  Andersen & Chen (1995)described this species based on a single female specimen from Jiangxi, China. In this study, we have collected both males and females from Hubeiand Zhejiang, China; thus, we can provide a description of the apterous male and additional distribution information. 4429869393 2022-07-11 NKUM Zhao-qi Leng & Zhen Ye China Ying-shan County 130 29.05711 Jiu-gong Mountain 1 107.09864 Huang-gang City 53 54 5 5 Hubei 4429869347 [318,1251,991,1020] 1919-06-02 IZAS China Nan-chang City 53 54 2 2 Jiangxi 4429869394 2015-07-29 NKUM Wen-bo Yi China Pan-an County 28.975029 Da-pan Mountain 1 120.52678 Jin-hua City 53 54 3 3 Zhejiang