Novius Koebelei Olliff in Craw, 1892: 14 Coquillett, 1893: 20 Lea, 1902: 493 Rodolia koebelei : Korschefsky, 1931: 101 Gordon, 1972: 26 Ślipiński, 2007: 143 Novius limbatus Blackburn, 1895: 254 Rodolia limbata : Ślipiński, 2007: 143 Rodolia blackburni Ukrainsky, 2009: 285 Novius limbata Blackburn, 1895 Novius tridens Lea, 1902: 492 Rodolia tridens : Ślipiński, 2007: 143 Novius simplicipennis Blackburn, 1895: 253 Rodolia simplicipennis : Ślipiński, 2007: 143 Novius discoidalis Blackburn, 1895: 253 Rodolia discoidalis : Ślipiński, 2007: 143 Novius tripustulatus Blackburn, 1895: 254 Rodolia tripustulata : Ślipiński, 2007: 143 Novius ruber Blackburn, 1889a: 148 Rodolia rubra : Ślipiński, 2007: 143 Revision Of The Australian Coccinellidae (Coleoptera). Genus Novius Mulsant Of Tribe Noviini Pang, Hong Tang, Xue-Fei Booth, Roger G. Vandenberg, Natalia Forrester, Juanita Mchugh, Joseph Ślipiński, Adam Annales Zoologici 2020 2020-03-30 70 1 1 24 Olliff in Craw Olliff in Craw 1892 [952,1308,180,203] Insecta Coccinellidae Novius GBIF Animalia Coleoptera 7 8 Arthropoda species koebelei  ( Figs 19–24, 48–59, 64, 76–80, 90, 96–99)      Novius KoebeleiOlliff in Craw, 1892: 14. TL: Australia, introduced to U.S.A.;  Coquillett, 1893: 20;  Lea, 1902: 493.     Rodolia koebelei: Korschefsky, 1931: 101;  Gordon, 1972: 26;  Ślipiński, 2007: 143.     Novius limbatus Blackburn, 1895: 254. TL: Queensland, near Cairns. Syn. nov.     Rodolia limbata: Ślipiński, 2007: 143.     Rodolia blackburni Ukrainsky, 2009: 285(replacement name for  Novius limbata Blackburn, 1895, not Motschulsky, 1866).     Novius tridens Lea, 1902: 492. TL: Queensland, near Cairns. Syn. nov.     Rodolia tridens: Ślipiński, 2007: 143.     Novius simplicipennis Blackburn, 1895: 253. TL: Queensland, Toowoomba. Syn. nov.     Rodolia simplicipennis: Ślipiński, 2007: 143.     Novius discoidalis Blackburn, 1895: 253. TL: Queensland, near Toowoomba. Syn. nov.     Rodolia discoidalis: Ślipiński, 2007: 143.     Novius tripustulatus Blackburn, 1895: 254. TL: Queensland, near Cairns. Syn. nov.     Rodolia tripustulata: Ślipiński, 2007: 143.     Novius ruber Blackburn, 1889a: 148. TL: New South Wales, Mulwala. Syn. nov.     Rodolia rubra: Ślipiński, 2007: 143.    Diagnosis.  Novius koebeleican only be diagnosed by the details of the male genitalia, in particular by the presence of the small apical barb at the apex of the penis guide.    Description.Length 2.7–3.5 mm. Body oval, widest near middle, 1.2–1.3 times longer than wide. Color pattern variable. Head, pronotum and scutellum usually uniformly dark. Elytra of typical form orange or red often with darker sutural stripe posteriorly and small lateral dark spot near midlength of elytron situated near lateral margin and continuing as darker stripe along lateral margin posteriorly. Melanic forms have elytra with black areas of various sizes or, rarely entirely black ( Figs 48–59, 64, 76–80, 90). Interocular distance in frontal view 1.6–2.0 times eye width. Male genitalia ( Figs 19–23, 108–111): parameres slender, not expanded apically; penis guide slightly longer than parameres, stout and narrowing apically with small apical barb. Penis bent and pointed apically. Female genitalia as in Figs 96–98.    Type material.  Novius KoebeleiOlliff in Craw: Lectotype, here designated, the specimen illustrated in Craw, 1892: frontispiece, plate 1, fig 3, “  Novius Koebelei, Olliff.: male enlarged.”;   Novius limbatusBlackburn: Lectotype, here designated ( BMNH), “ AustraliaBlackburn Coll. B.M. 1910-236/  Novius limbatusBlackb./ T 5938, Qu.”;   Novius tridensLea: Holotypemale: “10408  Novius tridensLea, N.S. Wales, TYPE, S.A. Museum”;   Novius simplicipennisBlackburn: Lectotype, here designated ( BMNH), “ AustraliaBlackburn Coll. B.M. 1910-236/  Novius simplicipennisBlackb./ T 4164, Qu”;   Novius discoidalisBlackburn: Lectotypemale, here designated ( BMNH), “ AustraliaBlackburn Coll. B.M. 1910-236/  Novius discoidalisBlackb./ T 5936, Qu.”;   Novius tripustulatusBlackburn: Lectotypefemale, here designated ( BMNH), “ AustraliaBlackburn Coll. B.M. 1910-236/  Novius tripustulatusBlackb./ T 5937, Qu.”;   Novius ruberBlackburn: Lectotypefemale, here designated ( BMNH), Blackburn: “ AustraliaBlackburn Coll. B.M. 1910- 236/  Novius ruberBlackb./ T 2963.”    Other specimens examined. New South Wales: Sydney, R.C.L. Perkins, 1942-95(4, BMNH); Exp.937, 8.9.59, Frank Wilson, Sydney, viii.1959, C.I.E. Coll. No. 16712, ANIC (5, ANIC).  Queensland:Tambourine Mts, 11-17.v.1935 (2, BMNH);  Brisbane, 28.28S 153.01E, ix.1992, V. Brancatini (2, BMNH);  MUS. VIC. ENT-1041, Goodna, x.1920, F.E. Wilson (1, VM);  19kmS of Bundaberg, Pine Ck, 9.v.1975, H. Frauca, (1, VM); Rockhampton, 10-11.iii.1965, Exp. Dr. J. Balogh, (1, ANIC); Caloundra, iv.1965, L. Simpson, captured on Acelyphasp. (4, ANIC);  Brisbane, 27.28S 153.01E, viii.1992, V. Brancatini, (LPL9521) Pred. of   Icerya seychellarumon   Livistona chinensis(1, ANIC); Kenmore, 27.30.7S 152.56.2E, 9.xi.1993, V. Brancatini,   Rodolia koebelei? LPL9532 (KE15), predator of   Icerya aegyptiacaon decoy  Ficus benjamina, (12, ANIC);  Brisbane, 27.28S 153.01E, 28.v.1992, V. Brancatini,   Rodoliasp? (LPL9521), Pred. of   Icerya seychellarumon   Livistona chinensis,(4, ANIC); Long Pocket, 27.30.6S 152.59.7E, v.1993, V. Brancatini,   Rodolia koebelei? LPL9532 (LP108), predator of   Icerya aegyptiacaon decoy  Ficus benjamina, (9, ANIC);  Indooroopilly, 27.30.0S 152.58.4E, 7.x.1994, O. Fahey & V. Branvatini,   Rodolia koebelei? LPL9533 (IP24), predator of   Icerya aegyptiacaon decoy  Ficus benjamina(6, ANIC);  Indooroopilly, 27.30.0S 152.58.4E, 1.vi.1994, V. Bran +catini & O. Fahey,   Rodolia koebelei? LPL9533 (IP31), predator of   Icerya aegyptiacaon   Livistona chinensis,(8, ANIC);  Indooroopilly, 27.30.0S 152.58.4E, 30.viii.1994, V. Branvatini,   Rodolia koebelei? LPL 9533 (IP22), predator of   Icerya aegyptiacaon   Livistona chinensis(5, ANIC); Kenmore, 27.37.3S 152.56.0E, ix.1994, V. Brancatini,   Rodolia koebelei,predator of   Icerya seychellarumon  Michelia figo(1, ANIC).  Northern Territory: DarwinNT, 12.27S 130.50E, x.1992, V. Brancatini (18, BMNH);  DarwinNT, 19.iv.1991, V. Brancatini (8, BMNH); Berrimah, Darwin, 16.iv.1991, V. Brancatini (2, BMNH); Alawa, Darwin, 22.iv.1991, V. Brancatini (6, BMNH);  Darwin, 19.iv.1991, V. Brancatini,   Rodoliasp. (LPL9507), ex   Icerya aegyptiacaon  Nandina domestica, Brit. Mus. (M.H.), 1993-94(6, BMNH);  12.46S 132.39E, 12kmNNW of Mt. Cahill, 25.x.1972, at light, E. Britton (1, ANIC);  Darwin, 12.27S 130.50E, x.1992, V. Brancatini,   Rodoliasp. (LPL9507), Lab. Cult. Reared on   Icerya aegyptiacaon  Ficus benjamina(4, ANIC);  Darwin, 12.27S 130.55E, x./ xi. 1993, V. Brancatini, LPL 9507, Laboratory culture reared on   Icerya aegyptiacaon  Ficus benjamina(45, ANIC).    Distribution. Australia( Fig. 24), introduced to the United States( California) and subsequently to many countries worldwide.    Remarks.  Novius koebeleihas been described or mentioned many times in the literature, usually with reference to Olliff (1895) who was originally cited as the author of this species. Coquillett (1893)had previously described the larval and egg stages of this species, and was subsequently cited as the author by Gordon (1972, 1985). Gordon (1972)designated a fourth instar larva as the neotypeof  Novius koebeleiCoquillettbut this action is invalid (based on an individual that is insufficient to ensure recognition), and unnecessary because the first description of this species appeared in 1892 inthe report by A. Craw who illustrated the adult and larvae accompanied by the name  Novius Koebelei, Olliff. Accordingto the ICZN Art 74.4, we here designate the specimen illustrated in fig. 3 of that paper as the lectotypeof  Novius koebeleiOlliff in Craw (1892). 2609449676 BMNH Australia Australia 8 9 T 5938 1 lectotype 2609449691 TYPE United Kingdom Wales 8 9 1 holotype 2609449648 BMNH Australia Australia 8 9 T 4164 1 lectotype 2609449660 BMNH Australia Australia 8 9 T 5936 1 lectotype 2609449603 BMNH Australia Australia 8 9 T 5937 1 lectotype 2609449632 BMNH Australia Blackburn 8 9 T 2963 1 lectotype 2609449635 1942 United Kingdom 8 9 2609449656 1935-05-11 United Kingdom 9 10 8 9 Queensland 2609449707 1992-09 United Kingdom -28.28 153.01 9 10 Brisbane 2609449686 1920-10 United Kingdom 9 10 Mus 2609449669 1975-05-09 United Kingdom 9 10 Mus 2609449681 1965-03-10 United Kingdom 9 10 Mus 2609449683 1965-04 United Kingdom 9 10 Mus 2609449619 1992-08 United Kingdom -27.28 153.01 9 10 Brisbane 2609449689 1993-11-09 United Kingdom 9 10 Rodolia Koebelei 2609449638 1992-05-28 United Kingdom -27.28 153.01 9 10 Brisbane 2609449703 1993-05 United Kingdom 9 10 Rodolia Koebelei 2609449612 1994-10-07 Iceland 9 10 Indooroopilly 2609449644 1994-06-01 Iceland 9 10 Indooroopilly 2609449657 1994-08-30 Iceland 9 10 Indooroopilly 2609449663 1992-10 Iceland -12.27 130.5 9 10 Rodolia Koebelei 2609449609 [816,1440,517,539] 1991-04-19 Iceland 9 10 Darwin 2609449702 [816,1440,548,570] 1991-04-16 Iceland 9 10 Darwin 2609449714 [816,1440,578,600] 1991-04-22 Iceland 9 10 Darwin 2609449668 1991-04-19 Iceland 9 10 Darwin 2609449677 1972-10-25 Iceland -12.46 132.39 9 10 Darwin 2609449601 1992-10 Iceland -12.27 130.5 9 10 Darwin 2609449671 1993-11 Iceland 10 11 -12.27 130.55 9 10 Darwin