Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae) Jałoszyński, Paweł Zootaxa 2017 2017-11-15 4349 1 1 120 SFFZ Jałoszyński, 2017 Jałoszyński 2017 [151,671,1724,1750] Insecta Staphylinidae Cephennodes GBIF Animalia Coleoptera 76 77 Arthropoda species gokarnaensis sp. nov. Cephennodes     Typematerial. Holotype: NEPAL( Kathmandu District): ♂, one label: " NEPAL: Bagmati/ Gokana For.  1400m/ nr. Kathmandu,1.4. / Löbl - Smetana81" [white, printed] ( MHNG).   Paratypes(5 exx): 4 ♀♀, same data as for holotype;  1 ♀, " NEPAL: Bagmati/ Gokana For.  1400m/ nr. Kathmandu,30.3. / Löbl - Smetana 81" [white, printed] ( paratypesin MHNGand cPJ).   Diagnosis.BL 1.13–1.20 mm; species of the  C. gokarnaensisgroup with apex of median lobe of aedeagus broadly rounded; apical projection of aedeagus directed outward from apex of median lobe.   Description.Body of male ( Fig. 231) dark brown, moderately stout, with relatively deep constriction between pronotum and elytra, strongly convex, covered with light brown setae; BL 1.13 mm. Head subtrapezoidal, HL 0.15 mm, HW 0.30 mm; vertex and frons confluent and weakly, evenly convex, supraantennal tubercles small, weakly elevated; eyes moderately large, strongly convex, coarsely faceted. Punctures on vertex and frons very small but distinct, sharply marked, slightly sparser on median area than on sides, separated by spaces subequal to diameters of punctures; setae sparse, short, suberect. Antennae moderately long and moderately strongly thickened, AnL 0.63 mm, pentamerous club indistinctly delimited; antennomere I about 1.2 × as long as broad; II narrower than I but similar in length, 1.3 × as long as broad; III slightly narrower and much shorter than II, about as long as broad, IV–VI each as broad as III but indistinctly longer, 1.1 × as long as broad, VII slightly broader and longer than VI, about 1.2 × as long as broad; VIII distinctly broader than VII and similar in length, about as long as broad; IX and X each distinctly larger than preceding ones, each about as long as broad; XI slightly broader than X, slightly shorter than IX–X together, about 1.6 × as long as broad.   FIGURES 223–224.Distribution of  Cephennodes(s. str.) letheanus  sp. n.(223, circle),  C.(s. str.) myagdiensis  sp. n.(223, triangle),  C.(s. str.) malla  sp. n.(224, circle),  C.(s. str.) gorkha  sp. n.(224, circle). Pronotum in dorsal view subrectangular with rounded sides, broadest near middle, PL 0.35 mm, PW 0.53 mm; anterior margin in strictly dorsal view slightly arcuate; lateral margins indistinctly microserrate, strongly rounded in anterior third, only slightly less so in posterior half and strongly convergent toward distinctly obtuse-angled hind corners; posterior margin shallowly bisinuate; lateral carinae distinct but narrow, fused with lateral margins; antebasal pits small and shallow, nearly equally distant from posterior and lateral pronotal margins. Punctures on disc slightly larger than those on frons and vertex, sharply marked, those in median area separated by spaces slightly shorter than diameters of punctures; setae moderately long, dense, suberect. Elytra elongate, oval, broadest slightly in front of middle, EL 0.63 mm, EW 0.58 mm, EI 1.09. Humeral calli weakly elevated; subhumeral lines very narrow, divergent posteriorly, equal to 0.45 EL, each developed as moderately sharp stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron located in shallow and round basal impression; apices of elytra separately rounded. Punctures slightly larger and slightly sparser than those on pronotum, relatively sharply marked, distinct; setae similar to those on pronotum, except for subapical region, where setae are distinctly denser and directed posteromesally. Hind wings present, long. Metaventrite with large, sharply delimited postmesocoxal impressions, with median area covered with fine, dense punctures. Abdomen and legs unmodified. Aedeagus ( Figs 233–234) stout, AeL 0.13 mm, apex of median lobe broadly rounded; apical projection broad, directed outward from apex of median lobe, rounded; parameres very slender, unequal in length, one not reaching and one exceeding apex of median lobe, each with one long apical seta.   FIGURES 225–226.Distribution of  Cephennodes(s. str.) tharepatianus  sp. n.(225, circle),  C.(s. str.) minisulcatus  sp. n.(225, triangle),  C.(s. str.) mustangensis  sp. n.(226, circle),  C.(s. str.) lalitpuranus  sp. n.(226, triangle).   FIGURES 227–228.Distribution of  Cephennodes(s. str.) paralalitpuranus  sp. n.(227, circle),  C.(s. str.) bahrabisensis  sp. n.(227, triangle),  C.(s. str.) bilaminatus  sp. n.(228, circle),  C.(s. str.) ghorepanianus  sp. n.(228, triangle). Female. Similar to male but with sparser setae on subapical region of elytra, with long wings; BL 1.15–1.20 mm(mean 1.17 mm); HL 0.15–0.16 mm(mean 0.15 mm), HW 0.29–0.30 mm(mean 0.30 mm), AnL 0.60–0.63 mm(mean 0.62 mm); PL 0.35–0.38 mm(mean 0.37 mm), PW 0.55–0.58 mm(mean 0.57 mm); EL 0.63–0.68 mm(mean 0.65 mm), EW 0.60–0.63 mm(mean 0.62 mm), EI 1.04–1.08.   Distribution( Fig. 245). Central Nepal.   Etymology.Locotypical, after Gokarna Forest and Gokarna village (today a part of Gokarneshwar municipality near Kathmandu).   Remarks.This species differs from similar  C. pseudogokarnaensisin more rounded pronotum and structure of the aedeagus. 1700659671 MHNG Nepal Kathmandu District 1400 Lobl - Smetana Kathmandu 76 77 1 1 Bagmati holotype 1700659639 MHNG Nepal Kathmandu District 1400 Lobl - Smetana Kathmandu 76 77 4 4 Bagmati paratype 1700659662 MHNG Nepal 1400 Bagmati 76 77 1 1 Bagmati paratype